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Publisher: Susan Winslow Senior Acquisitions Editor: Lauren SchultzAssociate Director of Marketing: Debbie ClareMarketing Assistant: Lindsay NeffDevelopmental Editor: Erica ChampionDevelopmental Editor: Irene PechDevelopmental Coordinator: Sara Ruth BlakeAssociate Media Editor: Allison MichaelSupplements Editor: Yassamine EbadatSenior Project Manager at Aptara: Sherrill ReddPhoto Editor: Christine BuesePhoto Researcher: Elyse ReiderArt Director: Diana BlumeText Designer: Marsha CohenIllustrations: ImagineeringIllustration Coordinator: Janice DonnolaProduction Coordinator: Lawrence GuerraComposition: Aptara%u00ae, Inc. Printing and Binding: RR DonnelleyLibrary of Congress Control Number: 2012950797North American EditionCover image: %u00a92009 Pfl icke and Sixt. Originally published in Th e Journal of Experimental Medicine. 206:2925-2935. doi:10.1084/jem.20091739. Image provided by Holger Pfl icke and Michael Sixt.International EditionCover design: Dirk KaufmanCover image: Nastco/iStockphoto.comNorth American EditionISBN-13: 978-14292-1919-8ISBN-10: 1-4292-1919-XInternational EditionISBN-13: 978-14641-3784-6ISBN-10: 1-4641-3784-6%u00a9 1992, 1994, 1997, 2000, 2003, 2007, 2013 by W. H. Freeman and CompanyAll rights reservedPrinted in the United States of AmericaFirst printingNorth American EditionW. H. Freeman and Company41 Madison AvenueNew York, NY 10010www.whfreeman.com International EditionMacmillan Higher EducationHoundmills, BasingstokeRG21 6XS, Englandwww.macmillanhighered.com/international