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Pseudochazara  mamurra (Herrich-Schäffer, 1846) Brown’s Grayling
                                                                                                                                                                                                          L1  Larva    L2 Larva and L1 moult.

                   A species of mountains, resembling P.  pelopea, and
                   overlooked until described from the Levant by the Turkish
                   researcher Ahmet Koçak (Koçak, 1978). Differs from P.
                   pelopea in the following characters:                                          Ļ  P. m. birgit
                   ‡  ,Q  P. mamurra  the hindwing margin is less serrated
                   than in P. pelopea.
                   ‡ 7KH ZLQJ IULQJHV DUH ZKLWH LQ P. mamurra and brown-                           P.  m. larseni                                                                                                            © Leah Benyamini
                   grey in P. pelopea.                                                       Ļ
                   ‡  $QGURFRQLDO  VFDOHV  LQ  P. mamurra are scattered                                                                                                                                                                                                                                                                                                                     8
                                                                                                                                                   Ļ
                   and the androconial patch is not well defined as in  P.                                                                        0HGLQDK
                   pelopea.  Under magnification, the androconial scales of                                                                                DGXOW                             © Leah Benyamini  © Dubi Benyamini  © Dubi Benyamini
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                   P. mamurra are 25% longer and three times wider than                                                                                   HJJ  1 1 1 1  1    2    3    4    5     6    7    8    9   10  1 1  1  1  1  1  12   Fully grown L2 Larva   Overwintrring L5,
                                                                                                                                                  lar v a
                   in P. pelopea.                                                          DGXOW  1    2    3    4    5    6    7    8    9   10  11  12           pupa                before moulting to L3  on Cyperus alternifolius 3 Feb. 2020
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                   ‡ P. mamurra flies mostly at elevations above 1750m.                lar v a
                                                                             pupa

                   P.  m.  larseni is rated (CR) critically endangered in the
                   Israeli Red List of butterflies (Renan et al., 2022 (in
                   prep.)).
                                                                                                                                      Israel, Beit-Arye, 300 m 26.8.1987  Israel, Fatsa’el, Lower Jordan Valley
                   %LRORJ\                                                                                                                                        -250 m (bsl) 16.10.1987
                                                                                                                                                             3
                   )OLJKW SHULRG  July to September
                   /LIH  KLVWRU\  univoltine. Early stages as reported from
                   Turkey: eggs are laid on the hostplant, are 0.9 mm in                                                                                                                     © Dubi Benyamini                © Dubi Benyamini
                   height, white, barrel-shaped with conical top and have 20                                                                   Jordan, Gawr as-Safi,  -360 m (bsl) 18.4.1998
                   vertical ribs. The larva hatches after about 10 days and                                                                                                             L5 after 3-4 overwintering months          2
                                                                                                                                                                                              6 Apr. 2020
                   has long white hairs, wide brown stripes on the sides and
                   a narrow reddish-brown dorsal stripe. The head is brown
                   and the 'tails', white. The larva enters diapause until the
                   spring thaw induces new growth of the hostplants. The
                   mature larva is brown with darker brown spots on the
                   sides; the sides of the body are light red-grey. The larvae                                                                                               © Dubi Benyamini
                   are nocturnal feeders, pupating below ground without the            P. m. larseni
                   use of silk. The pupa is 17 mm long. Adults eclose after a
                   month (Hesselbarth et al., 1995(2): 951).
                   5HFRUGHG KRVWSODQWV  Poaceae (Gramineae) - Poa annua.
                                                                                                            © Dubi Benyamini
                   'LVWULEXWLRQ
                                                                                        P.  m. larseni
                   TL: Mt. Ararat (Turkey). The species’ distribution ranges
                   from Greece to Caucasia, Armenia, Turkey, Syria, Lebanon,                                                                                                © Dubi Benyamini        © Dubi Benyamini         © Dubi Benyamini
                   Israel (Hermon), (absent from Cyprus, Jordan and Sinai),
                   Iran, Azerbaijan and Turkmenistan.                                                                                                                       Ex aestivating L5,      1
                                                                                                                                                                            18 July - 20 Oct. 2014
                   The Levant subspecies are:
                   Pseudochazara mamurra larseni  Koçak, 1978; TL: ?–
                   Syria, Lebanon and Israel (Hermon).
                   Pseudochazara mamurra brigit Gross, 1978. T.L. “Ziyaret
                   Geçidi, westlich Gürün, Province Sivas” - Antalya to the
                   northern fringes of the Levant in Mercin, Adana, Osmaniye
                   DQG *D]LDQWHS WR (OD]Lø
                                                                                                              © Stav Talal                       © Dubi Benyamini  © Dubi Benyamini           © Dubi Benyamini  © Dubi Benyamini  © Dubi Benyamini
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