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                      Melanargia syriaca (Oberthür, 1894)     Syrian Marbled White



                      A well-defined Levantine  Melanargia, more heavily
                      suffused with black than  M.  titea and  M.  wiskotti.
                      Flies in open glades in forests of oak and black pine
                      in typical ‘flip and glide’ flight similar to  Limenitis
 #
                      reducta. Males fly low among tall grasses in search
                      of females, rising to 1.5–2 m while overflying higher
                      vegetation. Based on genitalia and DNA analysis, the
                      revised status of the taxon  syriaca (as part of the                                                                           © Dubi Benyamini            © Dubi Benyamini                            © Dubi Benyamini
                      larger subspecific variation in the  larissa complex
 21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 5 - A | 22-01-02 | 12:34:17 | SR:-- | Magenta
 21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 5 - A | 22-01-02 | 12:34:17 | SR:-- | Yellow
                      recognized by Nazari  et al., (2009)), appears not to
 #21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 5 - A | 22-01-02 | 12:34:17 | SR:-- | Black
 21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 5 - A | 22-01-02 | 12:34:17 | SR:-- | Cyan
                      be supported by the morphological and biological
                      similarities to M. galathea.
                                                                                           DGXOW

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                                                                                          HJJ  1    2    3    4    5    6    7    8    9   10  11  12
                                                                                  lar v a
                                                                             pupa
                      %LRORJ\
                      )OLJKW  SHULRG  early May to late July, 700-1900 m
                      in Hatay (Atahan et al., 2018: 71); 2029-2119 m at
                      Al-Lazzab reserve Syrian Anti-Lebanon (Zarikian &
                      Ghrejyan, 2018).
                      /LIH KLVWRU\  univoltine. In the laboratory, a courting
                      male was seen to walk towards a female while
                      opening and closing its wings. Females show rejection
                      by flipping their wings quickly and raising the tip                                                                                                    © Dubi Benyamini                                © Dubi Benyamini
                      of the abdomen against the head of the male in a
                      previously unreported rejection posture. In Hatay,
                      females placed in a breeding cage with their favoured
                      blue flowers of  Scabiosa columbaria, laid eggs at
                      midday while fluttering and nectaring, and the eggs
                      fall to the ground. The spherical, white eggs are 1
                      mm in diameter, smooth, semi-glossy with a small
                      micropyle depression at the top. Eggs are completely
                      different from those of titea and wiskotti, but similar
                      to  M.  galathea. Under the microscope, 48 fine ribs
                      and a porous shell were observed (Hesselbarth  et
                      al., 1995(3): 485, DB pers. obs.). L1 hatches after
                      11 days, and does not eat the eggshell. The white L1
                      is 2.8-3 mm long, hairy and with a thin mid-dorsal
                      brown line and two sub-dorsal brown bands; the head
                      is wider than the body that tapers backwards. After
                      5-6 months of summer diapause, feeding commences
                      on young grasses, larval colour changing to green.
                      By the end of April/early May and feeding at night,                                                                                                    © Sylvain Cuvelier                              © Sylvain Cuvelier
                      L5 larvae grow to a chunky 25 mm long. The mature
                      larva is brownish-pink or light brown, has white short                                                                                              ©Ali Atahan Kütahya                             ©Ali Atahan Kütahya
                      hairs, four white longitudinal stripes, the upper two                                 © Dubi Benyamini
                      wider, with lower dark brown margins. Pupates on the
                      ground ventral side up without a silken web, beneath   'LVWULEXWLRQ
                      soft soil (Hesselbarth et al., 1995(3): 868) or possibly
                      among grasses. The pupa is 15 mm long, 6 mm wide,   TL: Akbès, Syrie, NE Hatay, S Turkey (as var. of galathea).
                      matt and light brown in colour, with a slightly coarse   'LVWULEXWHG  IURP  $GDQD   +DWD\   .DKUDPDQPDUDû  DQG
                      texture. Adults hatched after 12-14 days, males   NW Syria across CE Turkey and Hakkari Province (SE
                      emerging first.                                Turkey), possibly in N Iraq to NW Iran. In the Levant, the
                                                                     nominotypical ssp. is confined to C & S Amanos Mt and NW
                      5HFRUGHG  KRVWSODQWV  Poaceae (Gramineae); in the   Syria (Hesselbarth et al., 1995 (3): 767; Zarikian, 2016). It
                      laboratory, larvae preferred the annual grasses  Poa   overlaps slightly in its eastern limit with M. titea standfussi
                      annua - (illustr.) and  Brachypodium distachyon  over   in N Hatay and with M. t. titania in S Hatay (Atahan et al.,
                      perennial Piptatherum sp.                      2018: 71-72).
                                                                                                                                                                             © Sylvain Cuvelier                              © Sylvain Cuvelier
              76                                                                                                                                                                                                                          69
           2Butterflies of the Levant danaidae satyridae.indd   76                                             12/30/2021   4:23:32 PM  2Butterflies of the Levant danaidae satyridae.indd   69                                    12/30/2021   4:23:15 PM
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