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 Melanargia syriaca (Oberthür, 1894)     Syrian Marbled White



 A well-defined Levantine  Melanargia, more heavily
 suffused with black than  M.  titea and  M.  wiskotti.
 Flies in open glades in forests of oak and black pine
 in typical ‘flip and glide’ flight similar to  Limenitis
 #
 reducta. Males fly low among tall grasses in search
 of females, rising to 1.5–2 m while overflying higher
 vegetation. Based on genitalia and DNA analysis, the
 revised status of the taxon  syriaca (as part of the   © Dubi Benyamini  © Dubi Benyamini        © Dubi Benyamini
 larger subspecific variation in the  larissa complex
 21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 5 - A | 21-12-30 | 11:34:47 | SR:-- | Magenta
 21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 5 - A | 21-12-30 | 11:34:47 | SR:-- | Yellow
 recognized by Nazari  et al., (2009)), appears not to
 #21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 5 - A | 21-12-30 | 11:34:47 | SR:-- | Black
 21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 5 - A | 21-12-30 | 11:34:47 | SR:-- | Cyan
 be supported by the morphological and biological
 similarities to M. galathea.
                        DGXOW



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                       HJJ  1    2    3    4    5    6    7    8    9   10  11  12
               lar v a
          pupa
 %LRORJ\
 )OLJKW  SHULRG  early May to late July, 700-1900 m
 in Hatay (Atahan et al., 2018: 71); 2029-2119 m at
 Al-Lazzab reserve Syrian Anti-Lebanon (Zarikian &
 Ghrejyan, 2018).
 /LIH KLVWRU\  univoltine. In the laboratory, a courting
 male was seen to walk towards a female while
 opening and closing its wings. Females show rejection
 by flipping their wings quickly and raising the tip   © Dubi Benyamini                           © Dubi Benyamini
 of the abdomen against the head of the male in a
 previously unreported rejection posture. In Hatay,
 females placed in a breeding cage with their favoured
 blue flowers of  Scabiosa columbaria, laid eggs at
 midday while fluttering and nectaring, and the eggs
 fall to the ground. The spherical, white eggs are 1
 mm in diameter, smooth, semi-glossy with a small
 micropyle depression at the top. Eggs are completely
 different from those of titea and wiskotti, but similar
 to  M.  galathea. Under the microscope, 48 fine ribs
 and a porous shell were observed (Hesselbarth  et
 al., 1995(3): 485, DB pers. obs.). L1 hatches after
 11 days, and does not eat the eggshell. The white L1
 is 2.8-3 mm long, hairy and with a thin mid-dorsal
 brown line and two sub-dorsal brown bands; the head
 is wider than the body that tapers backwards. After
 5-6 months of summer diapause, feeding commences
 on young grasses, larval colour changing to green.
 By the end of April/early May and feeding at night,   © Sylvain Cuvelier                         © Sylvain Cuvelier
 L5 larvae grow to a chunky 25 mm long. The mature
 larva is brownish-pink or light brown, has white short   ©Ali Atahan Kütahya                  ©Ali Atahan Kütahya
 hairs, four white longitudinal stripes, the upper two   © Dubi Benyamini
 wider, with lower dark brown margins. Pupates on the
 ground ventral side up without a silken web, beneath   'LVWULEXWLRQ
 soft soil (Hesselbarth et al., 1995(3): 868) or possibly
 among grasses. The pupa is 15 mm long, 6 mm wide,   TL: Akbès, Syrie, NE Hatay, S Turkey (as var. of galathea).
 matt and light brown in colour, with a slightly coarse   'LVWULEXWHG  IURP  $GDQD   +DWD\   .DKUDPDQPDUDû  DQG
 texture. Adults hatched after 12-14 days, males   NW Syria across CE Turkey and Hakkari Province (SE
 emerging first.   Turkey), possibly in N Iraq to NW Iran. In the Levant, the
 nominotypical ssp. is confined to C & S Amanos Mt and NW
 5HFRUGHG  KRVWSODQWV  Poaceae (Gramineae); in the   Syria (Hesselbarth et al., 1995 (3): 767; Zarikian, 2016). It
 laboratory, larvae preferred the annual grasses  Poa   overlaps slightly in its eastern limit with M. titea standfussi
 annua - (illustr.) and  Brachypodium distachyon  over   in N Hatay and with M. t. titania in S Hatay (Atahan et al.,
 perennial Piptatherum sp.  2018: 71-72).
                                                  © Sylvain Cuvelier                              © Sylvain Cuvelier
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