Page 129 - Linear Models for the Prediction of Animal Breeding Values 3rd Edition
P. 129

(Continued )
                Effects                   Solutions

                Animals
                                        Direct effects       Maternal effects
                   1                        0.564                0.262
                   2                       −1.244               −1.583
                   3                        1.165                0.736
                   4                       −0.484                0.586
                   5                        0.630               −0.507
                   6                       −0.859                0.841
                   7                       −1.156                1.299
                   8                        1.917               −0.158
                   9                       −0.553                0.660
                  10                       −1.055               −0.153
                  11                        0.385                0.916
                  12                        0.863                0.442
                  13                       −2.980                0.093
                  14                        1.751                0.362
                Permanent environment
                   2                       −1.701
                   5                        0.415
                   6                        0.825
                   7                        0.461

            The solutions show little difference between the herds, but calves in pen 1 were
        heavier than those in pen 2 by about 6.85 kg at birth. The solution for level i of the
        fixed effect n can be calculated using Eqn 4.3 except that the sum of yields for the
        level of fixed effect is corrected in addition for maternal effects. That is:
                diag  in
                 ∑  y inf ∑ b ˆ inj ∑ a ˆ ink ∑ m ˆ inl ∑ pe ˆ  int t
                                               −
                                      −
                       −
                               −
            b ˆ  in  =   f=1  j  k       l       t                           (7.3)
                                 diag in
        where m  is the solution for level l of genetic maternal effects within level i of the nth
                inl
        fixed effect and all other terms are as defined in Eqn 4.3. Thus the solution for level 1
        of pen effect is:
                                    ˆ
                              ˆ
                         ˆ
                                                  ˆ
                                                      ˆ
                                          ˆ
                                              ˆ
            b ˆ  = [137 − (2hd  + hd  + hd ) − (a  + a  + a  + a )
             11            1    2    3    5   8   9   13
                     ˆ
                  − (2m  + mˆ  + mˆ ) − (2p ˆe  + p ˆe  + p ˆe )]/4
                      2   5    6       2    5    6
               = [137 − 4.82 − (−0.986) − (−2.832) − (−2.162)]/4
               = 34.540
               ˆ
        where hd  is the solution for level j of herd effect.
                j
            From the MME, the solutions for direct and maternal effects for animal i with
        progeny o are:
            ⎡  i u ˆ ⎤ ⎡  (         (       ⎤ −1   ⎡ u+ u ˆ  ⎤
                                                     ˆ
                                        1
                           1
                    1
                 =
            ⎢  ⎥ ⎢  n+ d + k )a 1    d + k )a  2 ⎥ ⎥  Hk 2  ⎢  s  d  ⎥
            ⎣  i ˆ m ⎦ ⎣  ( d+ k )a 2  n + d+ k )a  3 ⎦  ⎣m+ m ˆ  d ⎦
                                                     ˆ
                                    (
                                                      s
                                         1
                           1
                                 2
                   ⎡ y  − b ˆ  − m ˆ  − p ˆ  ⎤  ⎡ aˆ  − .(aˆ  )  ⎤
                                                  05
                 + ⎢  i  i   dam  dam ⎥ + H k  ⎢  o   mate  ⎥
                   ⎢ ⎣  y o  − b ˆ o  − u ˆ − ˆ p i ⎥ ⎦  3  ⎣ ˆ m o − .( ˆ m mate  ⎦ )  (7.4)
                                  −
                                                  05
                                o
        Maternal Trait Models: Animal and Reduced Animal Models              113
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