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CHAPTER 17  III The Leg  177


                                                                         the water’s surface (Figure 17-18). But the feet have
                                                                         another important function during takeoff: to push the
                                                                         bird’s torso off the water’s surface and greatly reduce
                                                                         energy-robbing friction, enabling the wings to lift the
                                                                         bird from the water. Once airborne and just like larger
                                                                         commercial aircraft, the feet are retracted and stowed
                                                                         beneath the undersurface of the torso or below the
                                                                         outstretched tail.
                                                                           When geese, pelicans, and cormorants land, they
                                                                         use their feet initially as air brakes, letting them dangle
                                                                         beneath their bodies as they begin to fl are  (Figure
                                                                         17-19). Just before touchdown, the feet are swung
                                                                         forward, tips upward, much like a pair of water skis
                                                                         immediately before the skier lands a jump (Figure
                                                                         17-20). Again, like commercial airliners, some landings
                                                                         are rougher than others, depending on the crosswinds
                                                                         and the roughness of the water. On touchdown, wing
                                                                         thrust is reversed and the feet are lowered into the
                                                                         water where they can function in a coordinated pad-
                                                                         dling fashion to advance, maintain position, or inde-
                                                                         pendently turn and maneuver.

                                                                         III THE VERSATILE PROPELLER:
                                                                             PROPULSION ON AND BENEATH
                                                                             THE WATER SURFACE

                                                                         To most observers, the function of a water bird’s feet
                    Figure 17-16 • An Amazon parrot perches on the hand of a   is clear: to propel it over the surface of the water (Figure
                    resident while being examined visually.
                                                                         17-21) or to dive (Figure 17-22). But what is not so
                                                                         readily apparent is how the feet of some water birds
                                                                         are used when submerged, especially predatory water
                    all in the same action. The inward convergence of the   birds in pursuit of prey (Figure 17-23). For example,
                    three backward-directed talons, combined with the    the cormorant is an exceptionally skilled underwater
                    forward-directed base claw, provides a near-inescap-  predator that uses its feet to both propel and guide
                    able, lethal grip that enables a raptor to carry away its   itself while chasing down small fi sh. Specifi cally, the
                    prey in either one or both feet (Figure 17-17).      cormorant’s feet can be angled horizontally to act as
                       During feeding, raptors often secure themselves   stabilizers or vertically to function as rudders, enabling
                    on a branch with one set of talons, while gripping   cormorants to change depth and direction almost
                    their prey with the other. The bird then feeds by    instantaneously.
                    tearing away the flesh and entrails of its victim with

                    its beak.
                                                                         Sandshoes
                                                                         The webbed feet of shorebirds serve as broad, fl exible
                    III THE ROLE OF THE FEET IN                          platforms on which to negotiate the sandy shoreline
                         TAKEOFF AND LANDING                             and its contiguous shallows. In principle, a shorebird’s
                                                                         feet function much like snowshoes, but unlike the rigid
                    During takeoff, most of the larger water birds, such as   winter counterparts, a shorebird’s feet are quite fl exi-
                    geese, use both their wings and feet to become       ble and are able to bend so as to contour irregular
                    airborne, with some, like the cormorant, requiring a   surfaces such as gravel, rocks, and vegetation. Sand-
                    longer “runway” than others. Ducks can become air-   shoes may also be conveniently collapsed during fl ight,
                    borne quite suddenly, especially if alarmed or startled,   which results in improved aerodynamics.
                    appearing to almost leap into the air, a kind of
                    jump-start.                                          Foot Injuries
                       When many water birds take off, they use their feet
                    in a manner that resembles running: initially getting   The most common serious foot injuries encountered in
                    the bird underway and then rapidly accelerating over   our practice are traumatic toe amputations, followed
















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