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The Chemistry and Fertility of Soils under Tropical Weeds   41



                              Some weeds are pioneer plants due to their fast growing characteristics and
                           their ability to propagate extensively with short life cycle.  Alang-alang (Imperata
                           cylindrica) is a very good example.  This weed may produce 44 new rhizomes of 13
                           m  long  with  as  many  as  646  tubers  in  18  weeks  of  growth  (Sriyani,  1993).    An
                           experiment in di Gunung Madu Plantation, Central Lampung, shows that a three
                           month  growth  of  on  tubers  of  teki  (Cyperus  rotundus)  in  the  absence  of  other
                           weeds  produced  as  many  as  524  new  tuber.    This  data  show  that  weeds
                           demonstrate the ability to quickly produce vegetative and generative organs such
                           as rhizomes or tubers.
                               The  biological  invasion  grows  with  modern  transportation,  global  trade,
                           transportation  of  germ  plasms  for  research  and  development,  biological
                           conservation, and also tourisms.  Biologists and weed scientists are now aware that
                           the spread of weeds is not only by winds or water, but  also by human.  Human
                           spread most beneficial plants from one area to others.  This activity may sometime
                           cause disasters.  For example, alang-alang was imported into the United States of
                           America  from  tropical  lands  for  land  conservation  to  cope  with  erosion  and
                           landslide problems but after years became a serious weed in some states of the
                           country like Florida and Louisiana (Tabor, 1949).  Kogan and Jepson (2007) argue
                           that biological invasion influence not only the local species and agroecosystem but
                           also the food safety and security as well as public health.  Acacia nilotica (Prickly
                           Acacia),  a  native  plant  of  the  subtropics  of  Africa,  Middle  East,  and  India,  was
                           brought to Australia for an esthetical plant and later for a  shading plant for animals
                           in pasture but later become weed in animal pasture  (Bolton, 1989).  This plant is
                           also  a  serious  weed  in  the  National  Park  of  Baluran,  East  Java,  while  Meremia
                           peltata  invaded  the  National  Park  of  South  Bukit  Barisan,  West  Lampung,  and
                           endangered  the  local  plants.    The  invasion  of  Acacia  nilotica  is  also  reported  to
                           disturb  the  ecosystem,  lower  the  existing  biodiversity,  and  endanger  the
                           population of wild bulls (Bos javanicus) (Siregar dan Tjitrosoedirdjo, 1999).
                              Most of plants introduced into Indonesia were important crops like rubber, oil
                           palm, cacao, soybean, corn, and cassava (Tjitrosoedirdjo et al., 2016).  However,
                           some plants introduced become invasive like  eceng gondok (Eichornia crassipes),
                           akasia  arabika,  kirinyuh  (Chromolaena  odorata),  and  sembung  rambat  (Mikania
                           micrantha).  Most of the 75 invasive plants reported by Tjitrosoedirdjo et al. (2016)
                           have  long  been  known  as  weeds  in  agricultural  lands  and  some  are  important
                           weeds  in  Indonesia  like  Imperata  cylindrica  (alang-alang),  Eichornia  crassipes
                           (eceng  gondok),  Mimosa  pudica  (putri  malu),  and  Mikania  micrantha  (sembung
                           sambat) shown in Fig. 3.2.  Some weeds also exist in non-agricultural lands called

                                                          Abdul Kadir Salam and Nanik Sriyani  – 2019
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