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and amphibians consume aquatic invertebrates and plants, and
                                                                          birds such as kingfishers, herons, and ospreys dine on fish and
                                                                          amphibians.


                                                                          Lakes and ponds are ecologically
                                                                          diverse systems

                                                                          Lakes and ponds are bodies of standing surface water. Their
                                                                          physical conditions and the types of life within them vary with
                                                                          depth and the distance from shore. As a result, scientists have
                     (a) Braided river in Nebraska                        described several zones typical of lakes and ponds (Figure 15.7).
                                                                             Around the nutrient-rich edges of a water body, the water
                                                                          is shallow enough that aquatic plants grow from the mud and
                                                                          reach above the water’s surface. This region, named the littoral
                                                                          zone, abounds in invertebrates—such as insect larvae, snails,
                                                                          and crayfish—on which fish, birds, turtles, and amphibians
                                                                          feed. The benthic zone extends along the bottom of the lake or
                                                                          pond, from shore to the deepest point. Many invertebrates live
                                                                          in the mud, feeding on detritus or on one another. In the open
                                                                          portion of a lake or pond, far from shore, sunlight penetrates
                                                                          shallow waters of the  limnetic zone. Because light enables
                                                                          photosynthesis (pp. 49–50), the limnetic zone supports phyto-
                                                                          plankton (algae, protists, and cyanobacteria; p. 93), which in
                                                                          turn support zooplankton (p. 93), both of which are eaten by
                                                                          fish. Within the limnetic zone, sunlight intensity (and there-
                                                                          fore water temperature) decreases with depth. Clear water
                                                                          allows sunlight to penetrate deeply, whereas turbid water
                     (b) Meandering river in Alaska
                                                                          (water with suspended solids) does not. Below the limnetic
                                                                          zone lies the profundal zone, the volume of open water that
                                                                          sunlight does not reach. This zone lacks photosynthetic life
                                                                          and is lower in dissolved oxygen than upper waters.
                                                                             Ponds and lakes change over time as streams and runoff
                                                                          bring them sediment and nutrients. Oligotrophic lakes and ponds,
                                                                          which are low in nutrients and high in oxygen, may slowly give
                                                                          way to the high nutrient, low oxygen conditions of eutrophic
                                                                          water bodies (jump ahead to see Figure 15.23, p. 430). Even-
                                                                          tually, water bodies may fill in completely by the process of
                                                                          aquatic succession (p. 106). As lakes or ponds change over time,
                                                                          species of fish, plants, and invertebrates adapted to oligotrophic
                                                                          conditions may give way to those that thrive in eutrophic condi-
                                                                          tions. These changes occur naturally, but eutrophication can also
                                                                          result from human-caused nutrient pollution (pp. 126–127).
                                                                             The largest lakes are sometimes known as inland seas.
                                                                          North America’s Great Lakes are prime examples. Lake Baikal
                                                                          in Asia is the world’s deepest lake, at 1637 m (just over 1 mile)
                     (c) Oxbow river in Arizona                           deep.  The Caspian Sea is the world’s largest body of fresh
                                                                          water, covering nearly as much area as Montana or California.
                     Figure 15.6  Rivers are classified according to their flow
                     across the landscape. The Platte River in Nebraska (a) shows an
                     example of a braided river, whereas the Wood River in Alaska   Freshwater wetlands include marshes,
                     (b) meanders across the landscape. The Horseshoe Bend portion   swamps, bogs, and vernal pools
                     of the Colorado River in Arizona (c) demonstrates an oxbow.
                                                                          Wetlands are systems in which the soil is saturated with water
                                                                          and which generally feature shallow standing water with
                        Rivers and streams host diverse ecological communi-  ample vegetation. There are many types of wetlands, and most
                     ties. Algae and detritus support many types of invertebrates,   are enormously rich and productive. In freshwater marshes,
                     from water beetles to crayfish. Insects as diverse as dragon-  shallow water allows plants such as cattails and bulrushes to
                     flies, mayflies, and mosquitoes develop as larvae in streams   grow above the water surface. Swamps also consist of shal-
             412     and rivers before maturing into adults that take to the air. Fish   low water rich in vegetation, but they occur in forested areas.







           M15_WITH7428_05_SE_C15.indd   412                                                                                    12/12/14   2:20 PM
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