366 - 7.8-8.2 - - - - (Lo , 2009 1,000-5,000 0.1-0.25 (Hydrogen partial pressure Nicotinamide adenine dinucleotide (NAD (Trace element Fe2+ Zn2+ Ni2+ Na+ Mg2+ Co2+ Fe-Fe hydrogenase Ni-Fe hydrogenase (Romero-Güiza , 2016 Lee 2001 Ferchichi 2005