Page 404 - Anatomy and Physiology of Farm Animals, 8th Edition
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Anatomy of the Digestive System / 389

               other structures. It is divided into a lesser   along  the  lateral  tongue  near  the  floor  of
                                                        the mouth. Numerous ducts pass directly
               omentum, extending from the lesser
  VetBooks.ir    curvature of the stomach to the liver, and   from the gland to open into the floor of the
                                                        mouth just ventrolateral to the tongue.
               the greater omentum, which is attached to
               the greater curvature of the stomach (and   With the exception of the horse, the sublin­
               the comparable portion of the ruminant   gual salivary gland also has a monostomatic
               stomach). The greater omentum spreads    portion that empties at the sublingual
               like an apron from the stomach to cover   caruncle on the floor of the mouth by way
               most of the ventral aspect of the mass of   of a single sublingual duct that runs parallel
               intestine.                               to the mandibular duct.
                                                           The salivary glands are classified as
               Accessory Digestive Organs               serous, mucous, or mixed. Serous glands
                                                        secrete a watery clear fluid, as compared
                                                        with mucous glands, which secrete mucus,
               In addition to the numerous small glands   a viscous material that acts as a protective
               located in the walls of the stomach and   covering for the surface of mucous mem­
               intestine, accessory glands include the sali­  branes. A mixed gland produces both
               vary glands, the pancreas, and the liver.  mucous and serous fluids. The parotid sali­
                                                        vary gland secretes primarily serous saliva;
               Salivary Glands                          mandibular and sublingual glands are clas­
                                                        sified as mixed glands in domestic farm
               The  salivary  glands of  domestic  farm   animals. Most of the minor salivary glands
                 animals comprise three pairs of well‐  have a mucous secretion.
               defined glands as well as scattered lobules
               of salivary tissue (the so‐called minor sali­
               vary glands). The chief salivary glands are   Pancreas
               the parotid, mandibular, and sublingual
               (Fig.  20‐18).  The  minor  salivary  glands   The  pancreas is a compound gland that
               include labial, buccal, lingual, and palatine   has both endocrine and exocrine portions.
               glands.                                  The exocrine portion of the pancreas pro­
                  The parotid salivary gland is located   duces sodium bicarbonate and digestive
               ventral to the ear in relation to the caudal   enzymes, which pass through the pancre­
               border of the mandible. In most animals,   atic ducts to empty into the duodenum
               the parotid salivary duct passes ventrad   close to the opening of the bile duct.
               and craniad on the medial side of the       The endocrine portion of the pancreas
                 caudal mandible then crosses the cheek   consists of isolated groups of pale‐staining
               superficially just cranial  to the masseter   cells scattered throughout the gland. These
               muscle. The duct then passes dorsad to   areas are called the pancreatic islets (for­
               penetrate the mucous membrane of the     merly islets of Langerhans). They produce
               oral vestibule near the third or fourth   the hormones that pass directly into the
                 maxillary cheek tooth.                 bloodstream (see Chapter 13), most nota­
                  The mandibular salivary gland is usu­  bly glucagon and insulin, which are the
               ally located ventral to the parotid gland,     primary regulators of blood sugar levels.
               just caudal to the mandible. The mandibu­   Grossly, the pancreas is an irregularly
               lar salivary duct passes forward along the   lobulated organ that lies adjacent to the
               medial side of the mandible to open ventral   proximal duodenum and frequently abuts
               to the tongue on the sublingual caruncle,   the stomach, the caudal vena  cava, and
               a small hillock of mucous membrane       caudal part of the liver as well. The pan­
               located on the floor of the mouth.       creas has the appearance of aggregated
                  The  sublingual salivary gland  is    nodules loosely united to form an elon­
               located deep to the oral mucous membrane   gated gland lying parallel to the duodenum.
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