Page 81 - Small Animal Clinical Nutrition 5th Edition
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Macronutrients         81


                  lulose, cellulose and lignin; however, the pectins are lost. Acid  Table 5-9. Characteristics of selected fiber ingredients.
        VetBooks.ir  detergent fiber (ADF) is determined by boiling the analytical  Fiber  Crude    fiber (%)*  Solubility  Fermentability
                                                                                          Total
                  sample in an acidic detergent solution. The fraction remaining
                                                                                         dietary
                                                                                   fiber
                                                                                   (%)*
                  contains cellulose and lignin. The result of subtracting the
                                                                                                  in water
                                                                       ingredient
                                                                                                              rate
                  amounts of ADF and NDF in a particular food or ingredient  Apple pectin  0  95  Soluble     Rapid
                                                                       Beet pulp    20     66    Insoluble  Moderate
                  approximates the hemicellulose content. Like the limitations of  Cellulose  80  98  Insoluble  Slow
                  crude fiber determinations, NDF and ADF do not measure the  Citrus pulp  12  77  Soluble    Rapid
                  more soluble fiber fractions of pectins and gums; therefore, they  Corn bran  19  90  Insoluble  Moderate
                                                                       Guar gum     0      81     Soluble     Rapid
                  are not widely used to determine fiber in pet foods.  Gum arabic  0      91     Soluble     Rapid
                                                                       Pea fiber    30     92    Insoluble  Moderate
                  Requirements                                         Peanut hulls  57    76    Insoluble    Slow
                                                                       Rice bran    44     13    Insoluble  Moderate
                  Fiber is not considered essential in the diets of cats and dogs,  Soy fiber  20  83  Soluble  Rapid
                  although it is often included in commercial foods. Overall, dogs  Soy hulls  34  69  Insoluble  Moderate/slow
                  and cats do not derive much energy from absorbing the typical  Sunflower hulls  54  80  Insoluble  Slow
                                                                       Wheat bran   10     43    Insoluble  Moderate
                  end products of bacterial fermentation; however, short-chain  Wheat middlings 7  46  Insoluble  Moderate
                  fatty acids are important in maintaining colonic health.  *As is basis.
                  Therefore, a small amount of fiber (<5%) that contains both
                  rapidly and slowly fermentable fibers is recommended in foods
                  for healthy pets (Chapters 13 and 20). Today, much interest  sources of fiber in dry extruded foods. Some pet foods contain
                  exists in human and veterinary nutrition about the role of  specific ingredients added to provide fiber. Fiber sources com-
                  “pharmacologic doses” of certain nutrients (e.g., fiber) in pre-  monly used in pet foods today include the hulls from rice, soy-
                  venting chronic diseases (Box 5-7).                 beans, peanuts and oats,dried beet pulp, various vegetable gums,
                    Fiber also aids in managing diseases such as obesity, diabetes  corn bran, wheat bran, rice bran, oat bran and more purified
                  mellitus, diarrhea, colitis and constipation. The types and  fiber sources such as oligosaccharides, cellulose and soy fiber.
                  amounts of dietary fiber required to assist in the management of  AAFCO publishes official definitions of fiber ingredients
                  these diseases can be found in Chapters 27, 29, 56, 62 and 64.  (2007). Table 5-9 lists common fiber ingredients with fiber con-
                                                                      tent and general classifications of solubility and fermentability.
                  Deficiencies
                  Total deficiency of fiber in typical pet foods is not a practical
                  problem because many ingredients contain some fiber.  PROTEIN/AMINO ACIDS
                  Homemade foods, veterinary therapeutic foods made with
                  more refined ingredients and purified diets used in research  Definition
                  studies can be extremely low in fiber. Some dietary fiber that  Proteins are large, complex molecules composed of hundreds to
                  produces short-chain fatty acids is usually recommended.  thousands of amino acids. Amino acids are composed of car-
                                                                      bon, hydrogen, oxygen, nitrogen and sometimes sulfur and
                  Excess/Toxicity                                     phosphorus. Four chemical groups covalently bonded to a car-
                                                                      bon atom form the general structure of amino acids. The four
                  Excess fiber may have undesirable effects. For instance, certain
                                                                      groups include a hydrogen atom, a carboxyl group (COOH),
                  fiber types decrease mineral absorption. The effects on mineral
                                                                      an α-amino group (NH ) and another chemical group specif-
                                                                                         2
                  absorption vary by type of fiber and the mineral. More rapidly
                                                                      ic for each amino acid (Table 5-10). Although hundreds of
                  fermentable fibers (e.g., pectins and guar gum) appear to  amino acids exist in nature, only 20 are commonly found as
                  decrease availability of some minerals, whereas fibers that con-  protein components. The amino acids found in mammalian
                  tain more cellulose have little effect on mineral absorption.  proteins are the L-isomer of α-amino acids, which means the
                    Excess fiber can dilute the energy and nutrient content of the  side chain unique to each amino acid is linked to the α-carbon
                  food to such an extent that an animal may have difficulty eat-  atom in the L (levorotatory) position.
                  ing enough of the food to meet its needs. Controlled levels of
                  fiber are advantageous in weight-reducing foods for dogs and  Structure
                  cats; however, such foods are fortified so that only total energy  Proteins are linear polymers of amino acids in which the amino
                  intake is low and other nutrients are present in adequate  group of one amino acid and the carboxyl group of another
                  amounts to meet daily requirements.                 amino acid are coupled together (peptide bond) (Figure 5-15).
                                                                      Amino acids arranged in chains are referred to as peptides.Two
                  Sources                                             bonded amino acids form a dipeptide, three a tripeptide and
                  The maximum crude fiber content of pet foods must be listed in  more than three a polypeptide.
                  the guaranteed analysis section of pet food labels in the United  Proteins can be described as having a primary, secondary, ter-
                  States. Most dog and cat foods have DM crude fiber contents  tiary or quaternary structure. The primary structure of proteins
                  of less than 5%. The fiber in most pet foods comes from a vari-  refers to the sequence (order) of amino acids along the polypep-
                  ety of ingredients. Grains such as whole corn and brown rice are  tide chain. The secondary structure of proteins refers to the
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