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Brooke RK, Cooper J, Shelton PA, Crawford RJM. 1982. Taxonomy, 2001. African Penguins as predators and prey – coping (or
distribution, population size, breeding and conservation of not) with change. South African Journal of Marine Science
the White-breasted Cormorant, Phalacrocorax carbo, on the 23: 435–447.
southern African coast. Le Gerfaut 72: 189–220. Crawford RJM, Dundee BL, Dyer BM, Klages NTW, Meÿer MA,
Brooke RK, Allan DG, Cooper J, Cyrus DP, Dean WRJ, Dyer Upfold L. 2007a. Trends in numbers of Cape Gannets
BM, Martin AP, Taylor RH. 1999. Breeding distribution, (Morus capensis), 1956/57–2005/06, with a consideration
population size and conservation of Grey-headed Gull Larus of the influence of food and other factors. ICES Journal of
cirrocephalus in southern Africa. Ostrich 70: 157–163. Marine Science 64: 169–177.
Cooper J. 1981. Biology of the Bank Cormorant, part 1: distribution, Crawford RJM, Dyer BM, Kemper J, Simmons RE, Upfold L. 2007b.
population size, movements and conservation. Ostrich 52: Trends in numbers of Cape Cormorants (Phalacrocorax
208–215. capensis) over a 50-year period, 1956–57 to 2006–07.
Cooper J, Brooke RK, Shelton PA, Crawford RJM. 1982. Emu 107: 253–261.
Distribution, population size and conservation of the Cape Crawford RJM, Cockroft AC, Dyer BM, Upfold L. 2008. Divergent
Cormorant Phalacrocorax capensis. Fisheries Bulletin South trends in Bank Cormorants Phalacrocorax neglectus
Africa 16: 121–143. breeding in South Africa’s Western Cape consistent with
Cooper J, Crawford RJM, Suter W, Williams AJ. 1990. a distributional shift of rock lobsters Jasus lalandii. African
Distribution, population size and conservation of the Swift Journal of Marine Science 30: 161–166.
Tern Sterna bergii in southern Africa. Ostrich 61: 56–65. Crawford RJM, Underhill LG, Altwegg R, Dyer BM, Upfold L.
Cooper J, Brooke RK, Cyrus DP, Martin AP, Taylor RH, Williams AJ. 2009. Trends in numbers of Kelp Gulls Larus dominicanus
1992. Distribution, population size and conservation of the off western South Africa, 1978–2007. Ostrich 80: 139–143.
Caspian Tern Sterna caspia in southern Africa. Ostrich 63: Crawford RJM, Altwegg R, Barham BJ, Barham PJ, Durant JM,
58–67. Dyer BM, Geldenhuys D, Makhado AB, Pichegru L, Ryan
Crawford RJM. 2009. A recent increase of swift terns Thal- PG, Underhill LG, Upfold L, Visagie J, Waller LJ, Whittington
asseus bergii off South Africa – the possible influence of PA. 2011. Collapse of South Africa’s penguins in the early
an altered abundance and distribution of prey. Progress in 21st century: a consideration of food availability. African
Oceanography 83: 398–403. Journal of Marine Science 33: 139–156.
Crawford RJM, Underhill LG. 2003. Aspects of breeding, molt, Crawford RJM, Dyer BM, Geldenhuys D, Makhado AB, Randall
measurements and population trend of Hartlaub’s Gull in RM, Upfold L, Visagie J, Waller L. 2012. Trends in numbers
Western Cape, South Africa. Waterbirds 26: 139–149. of crowned cormorants in South Africa, with information on
Crawford RJM, Cooper J, Shelton PA. 1982a. Distribution, diet. African Journal of Marine Science 34: 411–424.
population size, breeding and conservation of the Kelp Crawford RJM, Kemper J, Underhill LG. 2013a. African Penguin
Gull in southern Africa. Ostrich 53: 164–177. (Spheniscus demersus). In: Garcia Borboroglu P, Boersma
Crawford RJM, Shelton PA, Brooke RK, Cooper J. 1982b. PD (eds). Penguins Natural History and Conservation.
Taxonomy, distribution, population size and conservation University of Washington Press; Seattle and London: 211-
of the Crowned Cormorant Phalacrocorax coronatus. Le 231.
Gerfaut 72: 3–30. Crawford RJM, Randall RM, Whittington PA, Waller LJ, Dyer BM,
Crawford RJM, Shelton PA, Cooper J, Brooke RK. 1983. Allan DG, Fox C, Martin AP, Upfold L, Visagie J, Bachoo
Distribution, population size and conservation of the Cape S, Bowker M, Downs CT, Fox R, Huisamen J, Makhado
Gannet Morus capensis. South African Journal of Marine AB, Oosthuizen WH, Ryan PG, Taylor RH, Turpie JK. 2013b.
Science 1: 153–174. South Africa’s coastal-breeding white-breasted cormorants:
Crawford RJM, Williams AJ, Hofmeyr JH, Klages NTW, Randall population trends, breeding season and movements, and
RM, Cooper J, Dyer BM, Chesselet Y. 1995. Trends of diet. African Journal of Marine Science 35: 473–490.
African penguin Spheniscus demersus populations in the Crawford RJM, Randall RM, Cook TR, Ryan PG, Dyer BM, Fox R,
20th century. South African Journal of Marine Science 16: Geldenhuys D, Huisamen J, McGeorge C, Upfold L, Visagie
101–118. J, Waller LJ, Whittington PA, Wilke CG, Makhado AB. 2016.
Crawford RJM, Dyer BM, Cordes I, Williams AJ. 1999. Seasonal Cape cormorants decrease, move east and adapt foraging
pattern of breeding, population trend and conservation strategies following eastward displacement of their main
status of Bank cormorants Phalacrocorax neglectus off prey. African Journal of Marine Science 38: 373–383.
southwestern Africa. Biological Conservation 87: 49–58. Crawford RJM, Sydeman WJ, Thompson SA, Sherley RB,
Crawford RJM, David JHM, Shannon LJ, Kemper J, Klages NTW, Makhado AB. 2019. Food habits of an endangered
Roux J-P, Underhill LG, Ward VL, Williams AJ, Wolfaardt AC. seabird indicate recent poor availability of abundant forage
resources. ICES Journal of Marine Science 76: 1344–1352.
A view of some of the Stompneus Bay rocks with White-breasted Cormorants breeding in the foreground, 2015 (photo AB Makhado)
Sacred Ibis nesting at Velddrif Salt Works, 2018 (photo RJM Crawford)
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