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with reproductive cycles.  There was no apparent long-   the  animals  whose hunting  is prohibited  in  Angola.
                  term trend in annual values of densities of nests per km   Executive Decree No. 252/18 of 13 July 2018 – Angola’s
                  (Figure 4).                                       Red  List  of  Species,  lists the species of fauna and flora
                     Taking  into account the number of  nests recorded  at  that are at some risk or danger of extinction, which is why
                  localities regularly  monitored  in  the  Kitabanga  project  they enjoy protection. Executive Decree No. 469/15 of 13
                  area, the overall nesting area, that females may lay 1 to  July 2015 prohibits the slaughter, in national territory, of the
                  8 nests per season (Aguirre 2004) and intervals between   species listed in Annex I of the Convention on International
                  nesting of 2 or 3 years (Hirth 1997, Broderick et al. 2001),   Trade  in  Endangered  Species  of Wild  Fauna  and  Flora
                  the average annual nesting population of Green Turtles in   (CITES).
                  Angola during 2010–2021 was gauged to be of the order   In addition  to the Laws and Decrees, Dispatch
                  of 91 females, with a maximum of 136 and a minimum of  nº 1489/21 of 26 March 2021 restricts the occupation of, and
                  45 females. There was a slight increase but no significant   fishing activity in, the region of the mouth of the Longa River
                  trend over time in numbers of females nesting along the   in favour  of  the  conservation  of  sea turtles;  the  National
                  coast (Figure 5). It is important to note that nesting in con-   Strategy and  Action Plan for the Biodiversity (NBSAP)
                  secutive seasons (12-months apart) has been observed on   2007–2012  includes  the conservation  of sea turtles in
                  some occasions, but always at small percentages of the   actions A.1.3, A.2.5 and D.3.3; the Strategy and National
                  total breeding population  (Hirth 1997, Pilcher 2000). Us-  Action Plans for Biodiversity (NBSAP) 2019–2025 present
                  ing a remigration interval of 3 years (Seminoff et al. 2015),   recommendations for rescue of species affected by oil spill
                  the estimated number of nesting females in the population   accidents at sea with relevance to avifauna and sea turtles
                  was 135–408 animals. However, the numbers of animals   and mobilization of funds and carrying out detailed stud-
                  observed feeding at the mouth of the Cunene River have   ies on the current situation of endemic and/or endangered
                  been much higher (~ 500 animals per km, Morais 2013,   species such as sea turtles.
                  2015).                                              Angola  is a signatory to the Convention  on Biological
                                                                    Diversity (CBD), Convention  on International  Trade in
                  Olive Ridley Turtle (Lepidochelys olivacea)       Endangered  Species of  Wild Fauna and Flora (CITES),
                                                                    Bonn Convention  (CMS), and  Cartagena Protocol, so
                  Of  the sixteen permanent monitoring sites,  the highest   it is com- mitted to the protection  and  conservation  of
                  number of nests was consistently recorded at Longa, which   its  biological heritage,  including  sea  turtles. A  full  list of
                  had an average annual density of 155 nests/km (Figure 3).   Angolan laws, decrees and strategies, and of conventions
                  However, the Cuanza region, which was only sampled in a   to  which  Angola  is  a  party,  that  have relevance to  the
                  single season (2020/21), had a higher density of nests. The   conservation of its turtles is given in Appendix 1.
                  lowest average nesting density (9 nests/km) was at Dande   Sea turtles are listed on Angola’s  Red List of  species
                  (Figure 3). Across the 104 km of beaches monitored regu-   as: D. coriacea Endangered (EN, Category B); C. mydas,
                  larly, the average density of nests was 54/km (Figure 4).   C. caretta,  L. olivacea  and  E. imbricata  Vulnerable  (VU,
                  Extrapolation  of  the mean  density of  nests at  monitored   Category C) (Primo et al. 2022). In 2022, global populations
                  beaches to the entire 1,210 km of coastline where nest-   of these species were classified as: D. coriacea VU (Wallace
                  ing  occurred  indicated  that  on  average  60,783  nests   et al. 2013); C. mydas EN (Seminoff 2004); C. caretta VU
                  (range 30,391–91,174  nests) were laid annually  during   (Casale and Tucker 2017); L. olivacea VU (Abreu-Grobois
                  2010–2021,  suggesting laying  by an equivalent  number   and Plotkin 2008); and E. imbricata Critically Endangered
                  of females. Using a remigration  interval of 1.5 years   (Mortimer and Donnelly 2008). In the Southeast Atlantic,
                  (Miller 1997, Metcalfe et al. 2015), the estimated size of the   D. coriacea  was  regarded  to  be  Data  Deficient  because
                  breeding  population  was 45,586–136,761  females. How-   trend data were unavailable for Gabon, which hosted the
                  ever, the number breeding decreased over time (Figure 5).  majority of this subpopulation (IUCN 2013). In the South
                                                                    Atlantic, C. mydas was of Least Concern (Broderick and
                  Conservation status                               Patricio 2019).

                  Turtles in Angola are protected by law. Law nº 5/98 of 19  References
                  June  1998 –  Base Environmental  Law,  in its  article 13,
                  establishes measures  aimed  at  ensuring the  protection   Abreu-Grobois A, Plotkin P (IUCN SSC Marine Turtle Specialist
                  and conservation of biodiversity, including sea turtles. Law   Group). 2008. Lepidochelys olivacea. The IUCN Red List of
                                                                        Threatened Species 2008: e.T11534A3292503.
                  nº 6-A/04 of 8 October 2004 – Law on Biological Aquatic   Afonso EC.  1987.  Contribuição  para o Conhecimento  das
                  Resources,  prohibits possession, transport, storage,   Tartarugas  Marinhas  Cheloniidae  da  Baía  do  Mussulo.
                  processing  and  sale  of  species of  turtles listed in its    Relatório  de Estágio de Licenciatura em Biologia.
                  Annex I.  Law nº 6/17 of 24 of  January  2017  –  Law  on    Departamento de Biologia, Luanda, Angola.
                  Forests and  Wild Fauna, in its article  23 ensures  the   Aguirre JT.  2004. Estudio de la biologia de la reproduccion  de
                  protection  of migratory  species  of wild  fauna and their   las tortugas marinas del sur de la Isla de Bioko (Guinea
                  habitats. In its article  29, it prohibits  the harvesting,   Ecuatorial). Tesis Doctoral, Universitat de Valensia, Facultat
                  cutting, hunting and possession of  threatened forest  or   de  Ciències  Biològiques,  Departamento  de Zoologia,
                  fauna resources. Law nº 8/20 of 16 April 2000 – Law on   Valencia, 210 pp.
                  Environmental  Conservation  Areas, protects sea turtles   Bernardo J,  Plotkin PT.  2007.  An evolutionary  perspective on
                  and their habitats located in conservation areas.     the  arribada phenomenon and reproductive behavioral
                                                                        polymorphism  of olive ridley sea turtles (Lepidochelys
                     Likewise, several Decrees assume protection for  sea   olivacea).  In:  Plotkin PT  (ed.).  Biology and Conservation
                  turtles. The Joint Executive Decree nº 201/16 of 26 April   of  Ridley Sea  Turtles.  Johns Hopkins University Press,
                  2016  regulates hunting licenses and its  Table  2 shows    Baltimore, USA, pp. 59–87.


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