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Pseudochazara mamurra (Herrich-Schäffer, 1846) Brown’s Grayling
L1 Larva L2 Larva and L1 moult.

A species of mountains, resembling P. pelopea, and
overlooked until described from the Levant by the Turkish
researcher Ahmet Koçak (Koçak, 1978). Differs from P.
pelopea in the following characters: Ļ P. m. birgit
‡ ,Q P. mamurra the hindwing margin is less serrated
than in P. pelopea.
‡ 7KH ZLQJ IULQJHV DUH ZKLWH LQ P. mamurra and brown- P. m. larseni © Leah Benyamini
grey in P. pelopea. Ļ
‡ $QGURFRQLDO VFDOHV LQ P. mamurra are scattered 8
Ļ
and the androconial patch is not well defined as in P. 0HGLQDK
pelopea. Under magnification, the androconial scales of DGXOW © Leah Benyamini © Dubi Benyamini © Dubi Benyamini
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P. mamurra are 25% longer and three times wider than HJJ 1 1 1 1 1 2 3 4 5 6 7 8 9 10 1 1 1 12 Fully grown L2 Larva Overwintrring L5,
lar v a
in P. pelopea. DGXOW 1 2 3 4 5 6 7 8 9 10 11 12 pupa before moulting to L3 on Cyperus alternifolius 3 Feb. 2020
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‡ P. mamurra flies mostly at elevations above 1750m. lar v a
pupa

P. m. larseni is rated (CR) critically endangered in the
Israeli Red List of butterflies (Renan et al., 2022 (in
prep.)).
Israel, Beit-Arye, 300 m 26.8.1987 Israel, Fatsa’el, Lower Jordan Valley
%LRORJ\ -250 m (bsl) 16.10.1987
)OLJKW SHULRG July to September
/LIH KLVWRU\ univoltine. Early stages as reported from 3
Turkey: eggs are laid on the hostplant, are 0.9 mm in © Dubi Benyamini © Dubi Benyamini
height, white, barrel-shaped with conical top and have 20
vertical ribs. The larva hatches after about 10 days and Jordan, Gawr as-Safi, L5 after 3-4 overwintering months 2 2
6 Apr. 2020
has long white hairs, wide brown stripes on the sides and -360 m (bsl) 18.4.1998
a narrow reddish-brown dorsal stripe. The head is brown
and the 'tails', white. The larva enters diapause until the
spring thaw induces new growth of the hostplants. The
mature larva is brown with darker brown spots on the
sides; the sides of the body are light red-grey. The larvae © Dubi Benyamini
are nocturnal feeders, pupating below ground without the P. m. larseni Koçak, 1978
use of silk. The pupa is 17 mm long. Adults eclose after a
month (Hesselbarth et al., 1995(2): 951).
5HFRUGHG KRVWSODQWV Poaceae (Gramineae) - Poa annua.
© Dubi Benyamini
'LVWULEXWLRQ
P. m. larseni
TL: Mt. Ararat (Turkey). The species’ distribution ranges
from Greece to Caucasia, Armenia, Turkey, Syria, Lebanon, © Dubi Benyamini © Dubi Benyamini © Dubi Benyamini
Israel (Hermon), (absent from Cyprus, Jordan and Sinai),
Iran, Azerbaijan and Turkmenistan. Ex aestivating L5, 1
18 July - 20 Oct. 2014
The Levant subspecies are:
Pseudochazara mamurra larseni Koçak, 1978; TL:
"LIBANON, Antilibanon, Arsal" – Arsal, Lebanon. Syria,
Lebanon and Israel (Hermon).
Pseudochazara mamurra brigit Gross, 1978. T.L. “Ziyaret
Geçidi, westlich Gürün, Province Sivas” - Antalya to the
northern fringes of the Levant in Mercin, Adana, Osmaniye
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© Stav Talal © Dubi Benyamini © Dubi Benyamini © Dubi Benyamini © Dubi Benyamini © Dubi Benyamini
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2Butterflies of the Levant danaidae satyridae.indd 116 12/30/2021 4:25:35 PM 2Butterflies of the Levant danaidae satyridae.indd 125 12/30/2021 4:26:06 PM