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Satyrus ferula  (Fabricius, 1793)                                Great Sooty Satyr                               Pseudochazara lidya (Staudinger, 1878) Lydian Tawny Rockbrown



                    Mountain species that flies very actively in sparsely                                                            This little known local species is endemic to Turkey.
                    wooded, grassy slopes or rocky areas. Males patrol the                                                           Recorded in N Levant in Kilis and Adana provinces, S
                    biotope in search of females. The females are mostly                                                             Turkey (Atahan pers. comm. to DB). Flies in open Pinus
                    sedentary and spend long time resting in the shade of                                                            nigra and Quercus spp. woodland with sparse grasses
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                    vegetation, tending to fly for short distance. Mating takes                                                      and dry rocky areas. Adult’s emergence coincide with
                    place in the afternoon, soon after eclosing and only                                                             the blooming of  Onopordum spp. its preferred nectar
                    seldom afterwards. Adults take nectar from Centaurea,                                                            source. In the morning adults sit with open wings on
                    Onopordum and  Thymus spp. (Lafranchis  et al., 2015:                                                            bare ground or rock to warm up. Copulation takes place
                    704-705, Hesselbarth et al., 1995(2): 890).                                                                      in the afternoon (Hesselbarth et al., 1995(2): 946).
 21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 8 - B | 22-01-02 | 10:39:03 | SR:-- | Magenta
 21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 8 - B | 22-01-02 | 10:39:03 | SR:-- | Yellow
 #21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 8 - B | 22-01-02 | 10:39:03 | SR:-- | Black
 21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 8 - B | 22-01-02 | 10:39:03 | SR:-- | Cyan
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                                                                                                                                                                                                           HJJ  1    2    3    4    5    6    7    8    9   10  11  12
                                                                                                                                                                                                   lar v a
                    %LRORJ\                                                                                                          %LRORJ\                                                  pupa
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                                                                                          HJJ  1 1 1  2  2  3  3  4  5  6  6  7  7  8  9  91 0  1 1  1  1  1 2  2
                    )OLJKW SHULRG  June-July from 800 m to 2000 m in Hatay                 lar v a                                   )OLJKW SHULRG  June to August, from 850 m to 2200 m
                                                                             pupa
                    S Turkey (Atahan  et al., 2018: 89). DB observed it on                                                           (Hesselbarth et al., 1995(2): 946)
                    WKH XSSHU VORSHV RI 1XU 'DøODUL 0W  & $PDQRV UDQJH DW
                         P RQ    -XO\       KLOOWRSSLQJ"  DQG LQ .L]LOGDø 6                                                          /LIH KLVWRU\  univoltine. The egg is white with 20 vertical
                    Amanos range at 1550 m on 8 June 2013 and 26 June                                                                ribs. The 1.2 mm – long larva hatches after 11-12 days
                    2014 in open black pine forest.                                                                                  and doesn’t eat the eggshell. It is light brown, with dark
                                                                                                                                     mid-dorsal brown stripe and a wide, dark brown side
                    /LIH KLVWRU\  univoltine. Females prefer small LHP for egg                                                       stripe with black spots below. Overwinters as a young
                    laying. Eggs are laid singly on the hostplants leaves. The                                                       larva. The fully grown larva has a wide black unbroken
                    egg is 1.4-1.5 mm high and 1.1 mm wide, barrel-shaped                                                            mid-dorsal strip that is segmented in front part. The
                    with 12-14 vertical ribs. Hatching time may differ from a                                                        lateral line consists of a wide, somewhat curled band,
                    month in rainy areas to 7 months in dry areas. The larvae                                                        made  of alternating dark and lighter, almost rectangular
                    gnaw gradually through the eggshell that is thicker (20                                                          spots. The head is brown with dark “V” shape lines.
                    microns) compared to other satyrids (6-11 microns).                                                              Pupation is underground, pupa is reddish-brown. Adults
                    Young larvae (L1/L2) start diapause in late summer to                                                            eclose after 3-4 weeks (Hesselbarth  et al., 1995(2):
                    early winter. In lower altitude the larvae are slow growing                                                      945-946).
                    and feed in sunny days. From L3 the larvae feed at
                    night. L4-L5 larvae can wander a few meters in search                                                            5HFRUGHG KRVWSODQWV: Poaceae (Gramineae) spp.
                    of fresh hostplants. The mature larva has a pale cream
                    ground colour with dark stripes across the head capsule                                                                                                                              P. l. obscura
                    that extend backwards along the whole body. Pupation         Turkey, Hatay, S Amanus Mts, 1550 m.                'LVWULEXWLRQ
                    usually takes place in a subterranean chamber built by
                    the fully grown L5. The pupa is glossy brown, formed                                                             7/   ´'RUI  %R]GDø  LP  %R]  'DøODULLµ   3URYLQFH  ù]PLU   :
                    bottom up in early spring. The adult ecloses after 24-                                                           Turkey). This species is known only from W & S Turkey.
                    30 days. (Hesselbarth et al., 1995(2): 890; Lafranchis et                                                        One ssp. is recorded from the Levant: P.  l.  obscura
                    al., 2015: 704-705; SBN, 1987: 247).                                                                             (Staudinger, 1878), TL: “Kilikisher Taurus nördlich von
                                                                                                                                     Tarsus”.
                    5HFRUGHG KRVWSODQWV LQ (XURSH  Poaceae (Gramineae) –                                                                                                                                                     © Dubi Benyamini
                    Deschampsia caespitosa, Festuca & Stipa spp. (Tuzov et
                    al., 1997(1): 244-245).
                    'LVWULEXWLRQ
                    TL: “Italien, [Westalpen, Valdieri]”. Distribution ranges
                    from the Pyrenees to S Europe, Turkey, SW Russia and
                    E Ukraine eastwards to Kazakhstan, C Asia, Transbaikal
                    and W China. The nominotypical ssp. flies in N Levant;
                    in S Turkey and N Syria. S. f. ssp. hadjina Heyne, 1895,
                    TL: “Hadjin (Kleinasien), Mesopotamien” was described
                    from Adana, S Turkey but Hesselbarth et al., (1995) and
                    Sbordoni  et al., (2018) treated it as a synonym of the     7XUNH\  $QWDNLD  1XU 'DøODUL 0WV       P
                    nominotypical ssp.


                                                                                                                                                                               © Ali Atahan                                  © Leah Benyamini
                                                                                                            © Leah Benyamini
              122                                                                                                                                                                                                                        119




           2Butterflies of the Levant danaidae satyridae.indd   122                                            12/30/2021   4:25:57 PM  2Butterflies of the Levant danaidae satyridae.indd   119                                   12/30/2021   4:25:47 PM
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