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                    Maniola telmessia Zeller, 1847       Eastern Meadow Brown                                                                                                                                                                             21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 4 - A | 22-01-02 | 10:39:03 | SR:-- | Cyan



                    One of the commonest satyrids of the Levant, M. telmessia
                    resembles the European  M. jurtina with which it flies
                    together in S Turkey. Females aestivate in the summer                                                    4
                    months, their reappearance in autumn occasionally
                    leading to the belief that the species has several broods:
                    “..so on the coast (of Lebanon) at least, there are likely
                    to be three broods.” (Larsen, 1974: 138). A single                                                                         © Dubi Benyamini                   © Dubi Benyamini                           © Dubi Benyamini
                    observation of ravining-like behaviour was observed  in
                    Park Canada C Israel, on 1 April 2008 (Tomer, 2008b).
                    Territorial male expelling  V.  cardui out of its territory
                    was observed by DB near Belen, Hatay. Fruit nectaring
                    is seldom observed. Males eclosing several days before
                    the females. After mating, aestivating females hide                          DGXOW  1    2    3    4    5    6    7    8    9   10  11  12
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                    among rocks and bushes in shady places, flying shortly                 lar v a
                                                                             pupa
                    in morning or late afternoon for short nectaring, but
                    reappearing permanently to oviposit from September
                    until late October, seldom to early November. Rarely
                    single males that aestivated with females reappear
                    in the fall and are on alert should their “services”
                    be needed; on 27.9.2018 inside the Mediterranean                                                                             © Ofir Tomer                    © Dubi Benyamini                          © Christodoulos Makris
                    forest around Mitspe Hila in upper Galilee N Israel in
                    a six hours combined monitoring effort; 163 females
                    and one “stand by” sap-feeding male were observed
                    (Benyamini, 2018c). Similar phenomenon exist in C
                    Europe “..Regarding late flying  Maniola (jurtina males)                                                                                                                           © Dubi Benyaminiminini n n n n n n n n ni ni n ni n n n ni n n ni n n n n n n n n n n n n n n ni n n n n n n n n n n n n n n n n n n n n n n n n n n n n n n n n n i i i i i i i i i i i i
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                    from warm places in Austria I know the phenomenon and
                    it has always surprised me. Maybe a strategy to mate
                    with virgin aestivating females?” (Andrea Grill’s reply to
                    DB’s question re “stand by males”).
                    %LRORJ\
                    )OLJKW SHULRG  February (in upper Jordan Valley) to June,
                    Israel, from -100 m (bsl) at lower Nahal Amud, NW
                    Lake of Galilee (Caspi & Tomer, 2004) to 2000 m at Mt                                                                                              © Shalev Waismen          © Yuval Evron                 © Yuval Evron
                    Hermon local peak (Talal, 2009b); April–October SL to
                    2000 m in Hatay (Atahan et al., 2018: 95); 1545-1828
                    m Al-Lazzab reserve Syrian Anti-Lebanon range (Zarikian
                    & Ghrejyan, 2018); May-October SL to middle height in
                    Lebanon (Larsen, 1974: 138); March/April–September/
                    October from  -230 m to 1050 m, N Jordan (Katbeh-
                                                                                 n. ab. morag  Benyamini, 2021
                    Bader et al., 2003; Larsen & Nakamura, 1983).             Israel, Ma’ale HaHamisha, 800 m, 10.5.1980
                    /LIH KLVWRU\  univoltine. Females fly low, settle in open
                    places on litter, lower FW among RW and “disappear”,
                    or hiding under shrubs and their margins over the soil.
                    Egg laying is induced by rain; In Hatay, females that
                    were collected and caged from 21.9 until 25.9 along the
                    Amanos Mountains laid eggs simultaneously when rain
                    started and L1 started to hatch 7-10 days later on 2.10.
                    Females that were collected in various places on Mt
                    Carmel, N Israel between 27-29.9 started to lay eggs on
                    29.9 when rain started. In a cage that was located near
                    a western window with direct afternoon sun eggs were         n. ab. ofir  Benyamini, 2021
                    laid from 11:50 until 16:30 and first L1 were observed      Israel, Mt Matat, 800 m, 26.5.2009
                    five days later. Females prefer to lay in crevices of bark
                    of  Quercus spp and other Mediterranean wood like
                    Pinus, Pistacia and Ceratonia spp. But also on dry stalks
                    of grasses. Fresh egg is cream, turning to yellow a day
                    later and 2-3 days later gets reddish-brown spots. It is
                    barrel shaped 0.6 mm diam and 0.6-0.7 mm high. The
                    upper flat area is composed of an internal circle with                                 © Dubi Benyamini                                                © Samir Hamza                                     © Dubi Benyamini
                                                                                                                                                                                © Yuval Evron
                                                                                                                                                                                                                              © Dubi Benyamini
              64                                                                                                                                                                                                                          49
           2Butterflies of the Levant danaidae satyridae.indd   64                                             12/30/2021   4:23:00 PM  2Butterflies of the Levant danaidae satyridae.indd   49                                    12/30/2021   4:22:09 PM
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