Page 32 - Linear Models for the Prediction of Animal Breeding Values 3rd Edition
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Using similar arguments:
                                  2
                          2
            p  = var(y˜ ) = s  + 1/n(s )
             22      2    B       W
                2
                                                    2
         where s  is the between-cow variance and 1/n(s ) is the mean of the within-cow
                B                                   W
         variance. From Section 1.4:
              2  1  2
            s  =  s a
              B  4
         and for cow i in the group of five cows:
                          2
              2
            s  = var(y˜  − s )
             W       2i   B
              ˜
         where y  is the mean of the first two lactations for cow i. Since all five cows each have
               2i
         two records like Zena:
              2
                         2
                       1
            s  = (p  −  s a )
              W    11  4
         and:
                 2
                                2
            1/n(s ) = 1/n(p  −  s a )
                              1
                 W        11  4
         Therefore:
            p  =  s a  + 1/n(p  −  s a )
                               1
                                 2
                   2
                 1
             22  4         11  4
                          1
                 1
               = (289) + ( )(867 −  (289)) = 231.2
                                  1
                 4        5       4
            The index equations are:
                                − 1
            ⎡  1 b ⎤ ⎡ 867  72.25⎤ ⎡ 289  ⎤
                 =
            ⎢  ⎥ ⎢             ⎥ ⎢       ⎥
             b ⎣  2⎦ ⎣  72.25 231.2  ⎦ ⎣  72.25 ⎦
            The solutions are b  = 0.316 and b  = 0.213 and the index is:
                             1            2
            I = 0.316(230 − 250) + 0.213(300 − 250)
         The accuracy of the index is:
            r = ( é 0 316.  (289 ) + 0 213.  (72 5.  ) 289/  ) ù = 0 608.
                                              û
                 ë
         1.7.3  Prediction of aggregate genotype
         At times, the aim is not just to predict the breeding value of a single trait but that of
         a composite of several traits evaluated in economic terms. The aggregate breeding
         value (H) or merit for such several or m traits can be represented as:
            H = w a  + w a  + ...+ w a
                  1 1   2 2       m m
         where a  is the breeding value of the  ith trait and  w  the weighting factor, which
                i                                       i
         expresses the relative economic importance associated with the ith trait. The con-
         struction of an index to predict or improve H is based on the same principles as those
         discussed earlier except that it includes the relative economic weight for each trait.
            Thus:
                −1
            I = P Gw(y − m)                                                 (1.22)
          16                                                              Chapter 1
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