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(pp. 78, 98) or habitats (p. 79) within some specified area.   In some groups, many species evolved rapidly as popu-
                     Scientists may also consider the geographic arrangement of   lations spread across a variety of environments and adapted
                     habitats, communities, or ecosystems at the landscape level,   to local conditions. Other groups diversified because of a
                     including the sizes and shapes of patches and the connections   tendency to become subdivided by barriers that promote spe-
                     among them (pp. 131–133). Under any of these concepts, a   ciation (pp. 71–72).  Still other groups accumulated  species
                     seashore of beaches, forested cliffs, offshore coral reefs, and   through time because of low rates of extinction.
                     ocean waters would hold far more biodiversity than the same
                     acreage of a monocultural cornfield. A mountain slope whose
                     vegetation  changes  with  elevation  from  desert  to  forest  to   Many species await discovery
                     alpine meadow would hold more biodiversity than a flat area   Scientists often express biodiversity in terms of species rich-
                     the same size consisting of only desert, forest, or meadow.  ness because that component is most easily measured and is
                        The Serengeti’s open plains are vast, but the region holds   a good gauge for overall biodiversity. Yet we still are pro-
                     a diversity of habitats, including savanna (p. 115), grassland   foundly ignorant of the number of species that exist. So far,
                     (p. 113), hilly woodlands, seasonal wetlands, and rock out-  scientists have identified and described about 1.8 million spe-
                     croppings called kopjes. This habitat diversity contributes to   cies of plants, animals, and microorganisms. However, most
                     the rich diversity of species in the region.         of Earth’s species remain undiscovered. Estimates for the total
                                                                          number that actually exist range from 3 million to 100 million,
                     Some groups hold more species than others            with the most widely accepted estimates in the neighborhood
                                                                          of 14 million.
                     Species are not evenly distributed among taxonomic groups.   Our  knowledge  of  species  numbers  is  incomplete  for
                     In number of species, insects show a staggering predomi-  several reasons. First, many species are tiny and easily over-
                     nance over all other forms of life (Figure 11.3 and Figure 11.4).   looked.  These include bacteria, nematodes (roundworms),
                     Within insects, about 40% are beetles, and beetles alone out-  fungi, protists, and soil-dwelling arthropods. Second, many
                     number all non-insect animals and all plants. No wonder the   organisms are so difficult to identify that ones thought to
                     British biologist J.B.S. Haldane famously quipped that God   be identical sometimes turn out, once biologists look more
                     must have had “an inordinate fondness for beetles.”  closely, to be multiple species. This is frequently the case with



                                     Plants
                                     Plants



                      Fungi
                                                                           Insects






                                                                      Molluscs                       Figure 11.3 Some groups
                       Arachnids
                                                                       Crustaceans                   contain more species
                                               Birds                                                 than others.This illustration
                                                                                                     shows organisms scaled
                                                                                                     in size to the number of
                                                Bacteria                                             species known from each
                       Mammals                                                                       group, giving a visual sense
                                        Reptiles                         Protists                    of their disparity in species
                                                                                                     richness. Because most spe-
                                                                                                     cies are not yet discovered
                                                     Sponges      Flatworms                          or described, some groups
                          Amphibians     Archaea                                       Jellyfish     (such as bacteria, archaea,
                                                                                                     insects, nematodes, protists,
                                                                                                     and fungi) may contain far
                                                                                                     more species than we now
                      Annelids
                      Annelids                                                                       know. Data from Groombridge,
                                                                                        Fishes
                                                                                                     B., and M.D. Jenkins, 2002. Global
                                            Roundworms
                                            Roundworms         Echinoderms                           biodiversity: Earth’s living resources
                                                                                                     in the 21st century. UNEP-World
                                                                                                     Conservation Monitoring Centre.
                                                                                                     Cambridge, U.K.: Hoechst
             296                                                                                     Foundation.






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