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fuel reactions in the  2  Calvin cycle. In these reactions, carbon
                                                                          atoms from carbon dioxide are linked together to manufacture
                                                                          sugars. Photosynthesis is a complex process, but the overall
                                                                          reaction can be summarized with the following equation:
                                                                            6CO  1 12H O 1 the sun’s S C H O  1 6O  1 6H O
                                       Light energy                             2      2              6  12  6  2     2
                                                                                            energy     (sugar)
                                                                             The number preceding each molecular formula indi-
                                                         Chloroplast      cates how many of those molecules are involved in the reac-
                                                                          tion. Note that the sums of the numbers on each side of the
                     1  Light reactions  H O                      O 2     equation for each element are equal; that is, there are 6 C,
                                     2
                                                                          24 H, and 24 O atoms on each side. This illustrates how
                                                  ATP     ADP             chemical equations are balanced, with each atom recycled
                                                         NADP +           and matter conserved. No atoms are lost; they are simply
                                                 NADPH   Inorganic        rearranged among molecules. Note also that water appears
                                                         phosphate        on both sides of the equation. The reason is that for every
                                                                          12 water molecules that are input and split in the process,
                                                                          6 water molecules are newly created. We can streamline the
                                                                          photosynthesis equation by showing only the net loss of 6
                                                                          water molecules:
                       2  Calvin cycle  CO 2                     Sugars
                                                                             6CO  1 6H O 1 the sun’s energy S C H O  1 6O
                                                                                 2    2                     6  12  6   2
                                                                                                            (sugar)
                                                                             Thus, in photosynthesis, water, carbon dioxide, and
                                                                          light energy from the sun are transformed to produce sugar
                     Figure 2.12 In photosynthesis, autotrophs such as plants,
                     algae, and cyanobacteria use sunlight to convert water and     (glucose) and oxygen. To accomplish this, green plants draw
                     carbon dioxide into oxygen and sugar. In the light reactions,   up water from the ground through their roots, absorb carbon
                     water is converted to oxygen in the presence of sunlight, creat-  dioxide from the air through their leaves, and harness sunlight.
                     ing high-energy molecules (ATP and NADPH). These molecules   With these ingredients, they create sugars for their growth and
                     help drive reactions in the Calvin cycle, in which carbon dioxide is   maintenance, and they release oxygen as a by-product.
                     used to produce sugars. Molecules of ADP, NADP , and inorganic   Animals, in turn, depend on the sugars and oxygen from
                                                         +
                     phosphate created in the Calvin cycle, in turn, help power the light   photosynthesis. Animals survive by eating plants, or by eat-
                     reactions, creating an endless loop.                 ing animals that have eaten plants, and by taking in oxygen.
                                                                          In fact, it is thought that animals appeared on Earth’s surface
                                                                          only  after  the  planet’s  atmosphere  had  been  supplied  with
                        Some organisms use the sun’s radiation directly to pro-  oxygen by cyanobacteria, the earliest autotrophs.
                     duce their own food. Such organisms, called  autotrophs or
                     primary producers, include green plants, algae, and cyanobac-
                     teria (a type of bacteria named for their cyan, or blue-green,   Cellular respiration releases chemical energy
                     color). Autotrophs turn light energy from the sun into chemical   Organisms make use of the chemical energy created by pho-
                     energy in a process called photosynthesis (Figure 2.12). In pho-  tosynthesis in a process called cellular respiration, which is
                     tosynthesis, sunlight powers a series of chemical reactions that   vital to life. To release the chemical energy of glucose, cells
                     convert carbon dioxide and water into sugars,  transforming dif-  employ oxygen to convert glucose back into its original start-
                     fuse energy from the sun into concentrated energy the organ-  ing materials, water and carbon dioxide. The energy released
                     ism can use. Photosynthesis is an example of moving toward   during this process is used to form chemical bonds or to per-
                     a state of lower entropy, and as such, it requires a substantial   form other tasks within cells. The net equation for cellular res-
                     input of outside energy.
                                                                          piration is the exact opposite of that for photosynthesis:

                     Photosynthesis produces food for plants                     C H O  1 6O  S 6CO  1 6H O 1 energy
                                                                                                    2
                                                                                    12
                                                                                  6
                                                                                                          2
                                                                                            2
                                                                                       6
                                                                                  (sugar)
                     and animals
                                                                             However,  the  energy  released  per  glucose  molecule  in
                     Photosynthesis occurs within cell organelles called  chloro-  respiration is only two-thirds of the energy input per glucose
                     plasts, where the light-absorbing pigment chlorophyll (which is   molecule in photosynthesis—a prime example of the second
                     what makes plants green) uses solar energy (light energy from   law of thermodynamics. Cellular respiration is a continuous
                     the sun) to initiate a series of chemical reactions called  1  light   process occurring in all living things and is essential to life.
                     reactions. During these reactions, water molecules split and   Thus, it occurs in the autotrophs that create glucose and also
                     react to form hydrogen ions (H ) and molecular oxygen (O ),   in heterotrophs, organisms that gain their energy by feeding
                                              +
                                                                    2
                     thus creating the oxygen that we breathe. The light reactions   on other organisms. Heterotrophs include most animals, as
               50    also  produce small, high-energy  molecules that are  used  to   well as the fungi and microbes that decompose organic matter.



           M02_WITH7428_05_SE_C02.indd   50                                                                                     12/12/14   2:53 PM
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