Page 95 - Environment: The Science Behind the Stories
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from open water and depositing nutrients in feces that sink,
                                                                        they shift the community’s nutrient balance to the bottom and
                                                                        benefit species that feed there. Once they clear the water,
                                                                        sunlight penetrates more deeply, spurring the growth of large-
                                                                        leafed underwater plants and algae. In the long term, however,
                                                                        eutrophication (pp. 126–127, 430) may ensue, bringing harm
                                                                        to the system.
                                                                            In recent years, scientists have documented a surprising
                                                                        new twist: One invader is being displaced by another. The
                                                                        quagga mussel (Dreissena bugensis), a close relative of the
                                                                        zebra mussel from Ukraine, is spreading through the Great
                                                                        Lakes and beyond. This species, named after an extinct
                                                                        zebra-like animal, is replacing the zebra mussel in many
                     (a) Zebra mussels clog water intake pipes of power plants  locations.
                       and industrial facilities.                           In a second twist, today there are signs that the zebra
                                                                        mussel invasion may be losing steam. In a number of areas,
                                                                        zebra mussel populations have begun declining, including
                                                                        in part of Lake Huron, portions of the Mississippi River, the
                                                                        St. Croix River in Minnesota, Lake of the Ozarks in Missouri,
                                                                        and other locations. Some declines are due to displace-
                                                                        ment by quagga mussels, but for others the reasons are
                                                                        unknown. In some cases, predation by fish or ducks is
                                                                        being credited for driving down zebra mussel numbers, as
                                                                        native predators develop a taste for the invader. A similar
                                                                        process has already occurred in Europe, where waterfowl
                                                                        began preying heavily on zebra mussels and contributed to
                                                                        population crashes.
                                                                            In areas where zebra mussels are declining, some of
                                                                        the native fish and invertebrates that suffered from their
                                                                        arrival are now recovering. Ecologists are monitoring popu-
                     (b) Zebra mussels foul, starve, and suffocate native clams by  lations closely to see how the situation develops. No one
                       adhering to their shells and sealing them shut.
                                                                        expects zebra mussels to disappear or be eradicated, but
                     Figure 4.1 Invasive species can have severe economic and   there is now hope that some of their impacts might be
                     ecological impacts.                                reversed.




                     Species Interactions                                 Competition can occur when resources
                                                                          are limited
                     Interactions among species are the threads in the fabric of eco-
                     logical communities. By interacting with many species in a   When  multiple  organisms  seek  the  same  limited  resource,
                     variety of ways, zebra mussels have set in motion an array   their relationship is said to be one of competition. Compet-
                     of changes in the communities they have invaded. To under-  ing organisms do not usually fight with one another directly
                     stand why invasive species introduced by people can cause so   and physically. Competition  is generally more  subtle and
                     much disruption, we must first look at how species naturally   indirect, taking place as organisms vie with one another to
                     interact. Ecologists organize species interactions into several   procure resources. Such resources include food, water, space,
                     fundamental categories (Table 4.1).                  shelter, mates, sunlight, and more. Competitive interactions
                                                                          can take place between members of the same species, called
                                                                          intraspecific competition, or between members of different
                        Table 4.1   Species interactions: effects on Their   species, called interspecific competition.
                               Participants                                  If individuals of the same species are competing for lim-
                      Type of interaction   Effect on      Effect on      ited resources, then competition becomes more intense when
                                            Species 1      Species 2      there are more individuals per unit area (denser populations).
                                                                          This is density dependence (p. 86), and it can limit the growth
                      Mutualism                +              +
                                                                          of a population.
                      Predation, parasitism,   +              –              Whereas intraspecific competition is a population-level
                      herbivory
                                                                          phenomenon, interspecific competition affects communities.
                      Competition              –              –           Interspecific competition can give rise to different types of
               94     “+” denotes a positive effect; “–” denotes a negative effect.   outcomes. If one species is a very effective competitor, it may







           M04_WITH7428_05_SE_C04.indd   94                                                                                     12/12/14   2:55 PM
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