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                    1.  All butterflies in the colour plates are depicted in
                    their natural size, upperside (on left) and underside (on
                    right). The gender symbol is shown above each butterfly                    Distribution map                                          %XWWHUÁLHV
                    with the actual biotope for each taxon shown in the   Name of the Authority           Established
                    background.                                           and year of description  distribution
                    2. In this volume, each of the subfamilies is identified

                    by a different colour.                         Colour by   6FLHQWLÀF               Seasonal/
                                                                                               Temporary/Migrant
                    3. The distribution maps feature:              subfamily  name
                    - Permanent occupation, shown in red.                                              Common name                                 of the Levant
                    - Temporary incursions (migrations) or historic
                      occurrences are shown in faint red.             Pseudochazara mniszechii (Herrich-Schäffer, [1851])  Tawny Rockbrown  IUCN
                                                                                                           conservation
                    - A question mark indicates possible occurrence;   Pseudochazara mniszechii in the Levant is known only   category
                      usually used between known populations.   ? ?    from the Amanos Mts., S Turkey, were it is common. Fly   Behaviour
                                                                       in dry rocky areas  (Atahan et al., 2018: 86), on 24 June
                                                                            DW  .L]LOGDø       P 6 $PDQRV 0W LW ZDV Á\LQJ LQ RSHQ
                    - A dotted line marks anticipated expansion        FOHDULQJV RI EODFN SLQH IRUHVW  '%   2ÀU 7RPHU SHUV  REV      modules               and nearby areas
                                                                       UHSRUWHG E\  7X]RY             LQ ´GU\ FOD\H\ VWRQ\ VORSHVµ
                                                                       $GXOWV  WDNH  QHFWDU  IURP  WKH  ÁRZHUV  RI  Carduus nutans,
                      or possible presence.                            Echinops pungens,  Onopdrum anatolicum (Hesselbarth et
                                                                       al                  Centaurea VSS  (Asteraceae) and Silene
                                                                       compacta  &DU\RSK\OODFHDH                            adult  Life-cycle


                                                                                          1    2    3    4    5    6    7    8    9   10  11  12


                                                                                                  9

                                                                                                   1
                                                                                                  9

                                                                                                             egg
                    - Black arrows indicate small localities or point to  Biology                    lar v a  1    2    3    4    5     6  7  8  8     9   10  1    1  12  activity
                                                                                                pupa
                      the species’ occurrence beyond the range of the   Flight period:  -XQH     6HSWHPEHU   IURP        WR        P   periods.
                                                                       (Atahan et al.,              -XO\ $XJXVW IURP      WR      P
                      distribution map.                                 7X]RY
                                                                       Life history:  XQLYROWLQH   WKH  HJJV  DUH  ODLG  VLQJO\  DQG  DUH
                    Maps were based on data gleaned from literature    ZKLWH  EDUUHO VKDSHG ZLWK    YHUWLFDO ULEV  /  KDWFKHV DIWHU   black margins
                                                                            GD\V  FD   PP ORQJ  OLJKW EURZQ ZLWK VSDUVH ZKLWH KDLUV
                                                                       DQG D GDUN PLG GRUVDO VWULS  7KH ODUYDH IHHG DW QLJKW  'XULQJ
                    and butterfly collections (private and museum),    the day they hide at the base of the LHP. The mature larva   indicate aestivation
                                                                       LV  FKXQN\          PP  ORQJ   JUH\LVK EURZQ   ZLWK  PLG GRUVDO
                                                                       OLQH RI EODFN VHJPHQWV  VXE GRUVDO ZKLWLVK EDQGV ZLWK WLQ\
                    correspondence with professional lepidopterists and   UHGGLVK VKRUW VWULSV DQG ORQJLWXGLQDO VLGH EOXLVK JUH\ EURDG   or diapausing
                                                                       VWULSV  IRXU RI WKHP SURFHHG RQ WKH IRUHKHDG  7KH VKRUW WDLOV
                                                                                                         periods.
                                                                       DUH RII ZKLWH  3XSDWLRQ WDNHV SODFH D IHZ FP EHORZ VXUIDFH LQ
                    the authors’ data banks (DB and 4D MicroRobotics for   VRIW VRLO  7KH SXSD LV OLJKW UHGGLVK EURZQ  7KH DGXOW HFORVHG            Southern Turkey, Cyprus, Syria, Lebanon, Israel,
                                                                       after 4 weeks (Hesselbarth et al
                    the Eastern Mediterranean and Egypt, EJ for Cyprus).  Recorded hostplants: Poaceae (Gramineae) - Festuca ovina   Natural
                                                                       (Atahan et al., 2018: 86). Poa annua, P. pratensis (Tuzov,

                    4. Life-cycle periods are shown for adult, egg, larval   Distribution                   biotope                                 Jordan, Egypt, Sinai & North-West Saudi Arabia
                    and pupal stages. Dark green indicates known activity,   7/  ´9RQ 7RNDW LQ .OHLQDVLHQµ  $QDWROLD  7XUNH\   'LVWULEXWHG   ‹ 'XEL %HQ\DPLQL
                                                                       IURP  1   *UHHFH  WR  7XUNH\   6  *HRUJLD   $UPHQLD  DQG  1  ,UDQ
                                                                        1D]DUL                  DEVHQW  IURP  6\ULD   /HEDQRQ   ,VUDHO
                    light green estimated (or data from outside the Levant)   &\SUXV  -RUGDQ DQG 6LQDL   7KH QRPLQRW\SLFDO VVS  ÁLHV LQ WKH   Life-cycle
                                                                       Levant.
                    and dark green with black margins, diapausing/                       © Adam Warecki  © Adam Warecki  © Adam Warecki  stages and
                    aestivation periods. Our data is based upon (known)                                     hostplant(s)
                    past and present records but, as elsewhere in the
                                                                                            © Adam Warecki  © Adam Warecki
                    world, a changing climate may influence the timing of                                   male &
                    life-cycle events in the future; in short winters, adults                               female
                    may hatch weeks or even months ahead of their known                                     in nature
                    past records.                                                  © Adam Warecki  ‹ 2ILU 7RPHU  ‹ 0HKHPHW *ƺO
                    5. Despite major efforts by the author in attempting to   118
                    breed all butterflies found in the Levant, several are           Male          Female
                    rare, recent arrivals or pose problematic access to their
                    breeding areas. Nothing is presented for those species                                                                                                       Vol. III
                    where early stages are unknown (except F.P.), but on a
                    few occasions where early stages are known for another                                                                                                 1\PSKDOLGDH
                    subspecies (or a similar species) we use alternative   Upperside                      Underside
                    photographs, denoting these with a narrow green frame.                                                                             Libytheinae, Danainae, Chraxinae,
                    6. The sample page on right provides explanations for
                    titles and notations.                                                                                                            Satyrinae,  Limenitidinae, Bibilidinae,
                                                                                                                                                               Heliconiinae, Nympalinae,
                    $EEUHYLDWLRQV XVHG IRU KRVWSODQW SUHIHUHQFHV
                    [M] - main/preferred; [C] - cultivated; [S] - southern arid areas; [?] - possible hostplant; [L] - captivity, lab.
                    ,QWHUQDWLRQDO 8QLRQ IRU &RQVHUYDWLRQ RI 1DWXUH  ,8&1  5HG /LVW &DWHJRULHV
                    In this series of books (Vols. I-IV), we adopt the IUCN listings of
                    conservation categories as defined in van Swaay et al. (2010).   Regionally Extinct  Least Concern
                    Of 11 notations, we use nine including:
                    ‘'Regionally Extinct’ (RE), ‘Critically Endangered’ (CR),   Critically Endangered  Data Deficient
                    ‘Endangered’ (EN), ‘Vulnerable’ (VU), ‘Near Threatened’ (NT),                                                                                       Dubi Benyamini
                    ‘Least Concern’ (LC), ‘Data Deficient’ (DD), ‘Not Applicable’   Endangered        Not Applicable
                    (NA) and ‘Not Evaluated’ (NE).
                                                                                                                                                      English editors: Eddie John & Arthur M. Shapiro
                    We use  categories as defined in Numa et al. (2016) for those   Vulnerable        Not Evaluated
                    Mediterranean butterflies found in the Levant, showing these in
                    black, e.g. ‘LC’. However, where the conservation status in the                   Least Concern                                                    Design Leah Benyamini
                                                                                                      (Mediterranean
                    Levant (specifically Israel) differs, we follow that given in the   Near Threatened  assessment)
                    Israeli Red List Assessment (in prep.) denoting this in green,   Mediterranean assessment (Numa et al., 2016) – in black lettering.
                    the colour used also for non-Mediterranean species, e.g. ‘LC’.   Levant (Israeli Red Book) assessment – in green lettering.
              14
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           2Butterflies of the Levant danaidae satyridae.indd   14                                             12/30/2021   4:20:16 PM  2Butterflies of the Levant danaidae satyridae.indd   3                                     12/30/2021   4:20:06 PM
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