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Subfamily Charaxinae                                 Charaxes hansali                                             Melanitis leda (Linnaeus, 1758)       Common Evening Brown

                                                                        C. & R. Felder, [1867]

                                                                        Cream-banded Charaxes                                        A rare species in the Levant’s southern fringes. This large and
                                                                                                                                     unmistakable Satyrid is unique among the region butterflies;
                                                                                                                                     being a crepuscular species, flying mainly at dusk, hence its
                                                                        This powerful flyer is one of the largest butterflies of     common name. Adults may vary greatly in the wings’ shape and
 #
                                                                        the region, rivalling in size its related species C. Jasius.   underside coloration between wet and dry seasons, with the first
                                                                        Better adapted to arid areas than other congeners;           having more angular wings and much reduced and often absent
                                                                        inhabiting desert oasis and scrap-vegetation areas           eye spots.
                                                                        supporting its LHP. Males are territorial. Adults are
                                                                        attracted to fermenting fruits and feed on sap.
 21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 3 - B | 22-01-02 | 12:34:17 | SR:-- | Magenta
 21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 3 - B | 22-01-02 | 12:34:17 | SR:-- | Yellow
                                                                                                                                     %LRORJ\
 #21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 3 - B | 22-01-02 | 12:34:17 | SR:-- | Black
 21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 3 - B | 22-01-02 | 12:34:17 | SR:-- | Cyan
                                                                        Biology                                                      )OLJKW  SHULRG  In Saudi Arabia reported in May (Wadi Dur, T.
                                                                                                                                     Larsen, 1980; A.R. Pittaway, 1981), October (Wadi Sharas,                         DGXOW  1      2      3      4      5      6      7      8      9       1 0      1  12


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                                                                                                                                                                                                               HJJJ
                                                                        Flight period: recorded in Gebel Elba, SE Egypt in March     T. Larsen, 1981) and December (Dhofar, T. Michels, 1979).                lar v a
                                                                                                                                                                                                  pupa
                                                                        and April (Gilbert & Zalat, 2007: 101) ~1000-1800 m.         Elsewhere fly year round (Larsen, 1983).
                                                                        Recorded in Jebel el Lawz, Hejaz, NW Saudi Arabia in
                                                                        April, 2000-2500 m. In Oman recorded in March, 400-          /LIH KLVWRU\   not reported from the Levant. A fully grown 40 mm
                                                                        1000 m (Larsen, 1983: 419). Elsewhere year round.            long larva was found in the Lemur’s Anja Community Reserve in
                                                                                                                                     SW Madagascar in mid-April 2018 and was bred to male adult
                                                                        Life history: not reported in the Levant. In Africa: the     by DB (illustr.). The larva was light green with thin dark green
                                                                        singly laid egg is spherical with ridged flat top, cream     dorsal line and three wider lines along both sides, the lower one
                                                                        and change to yellow, a brown ring appears later and         being the wider. The body was covered with small white setae
                                                                        the egg becomes brown before L1 hatches after 7-10           and short white hairs. The body ends with whitish-green fork.
                                                                        days. The newly hatched larva it olive green with black      The head had two “horns”, dark green, with dark brown strip
                                                                        head. The mature larva resembles that of C. Jasius: is       runs up its sides to the “horns”, bordering a thinner white strip.
                                                                        55 mm long, green with thin yellow strips on both sides      The head was covered with long hairs, white on the green and
                                                                        and a mid-dorsal small oval reddish-purple spot. The         white areas, dark brown and white on the brown areas and horns       dry season form
                                                                        pupa is suspended from the LHP, 28 mm long, green            (illustr.). The green pendulous 17 mm long pupa is typical Satyrid
                                                                        and resembles that of C. Jasius. The adult eclosed after     shape, it remained green until one day before hatching in the
                                                                        3 weeks (Henning, 1988: 96).                                 lab. after eight days.
                                                                        Recorded hostplants: Not reported in the Levant,             5HFRUGHG KRVWSODQWV  Poaceae (Gramineae).
                                                                        elsewhere  Salvadora persica or  Dobera glabra
                                                                        (Salvadoraceae). Also known to feed on Osyris (illustr.)
                                                                        and Colpoon spp. (Santalaceae), but these plants are         'LVWULEXWLRQ
                                                                        extremely rare in NW Arabia (Collenette, 1999: 666).
                                                                                                                                     TL: “China”.  This species is widely distributed throughout all
                                                                                                                                     the Palaeotropical and Australian regions with numerous sspp.
                                                                        Distribution                                                 described throughout its range. The Levant’s specimens belong
                                                                                                                                     to the nominotypical ssp.
                                                                        TL: “Africa, Bogos” (N Ethiopia). Distributed from E                                                                              wet season form
                                                                        Africa, Tanzania and Kenya, to SE Egypt and the Arabian
                                                                        Peninsula. Three subspecies were reported in the wider
                                                                        Levant: C. h. hansali C. & R. Felder [1867] - confined
                                                                        to southeast Egypt in Gebel Elba.  C.  h. arabica Riley,
                                                                        1931. TL: “S. E. Arabia, Qara Mts and Wadi Balsh” -
                                                                        recorded from Dhofar & Oman (Larsen, 1983: 418)                                                                                                      © Dubi Benyamini
                                                                        and C. h. yemeni Turlin, 1998, described from Yemen.
                                                                        In April 2001 it was discovered at Jebel el Lawz, NW
                                                                        Hejaz, a northwards expansion of 950 km from its
                                                                        known record in Taif (Pittaway, 1985), (Benyamini &
                                                                        Müller, 2020).                                                                                   © Dubi Benyamini


                                                                                                                                                                                                                             © Dubi Benyamini




                ©  Limor Gefen                       Charaxes jasius                              © Hilary Welch                                       © Leah Benyamini         © Leah Benyamini         © Dubi Benyamini    © Dubi Benyamini







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           2Butterflies of the Levant danaidae satyridae.indd   38                                              1/2/2022   12:14:28 PM  2Butterflies of the Levant danaidae satyridae.indd   43                                    12/30/2021   4:21:51 PM
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