Page 164 - Linear Models for the Prediction of Animal Breeding Values 3rd Edition
P. 164

DIM
                           0                 1                2                 3
                          F                F                 F                F
          4              1.000            0.000             0.000            0.000
         38              0.333            0.667             0.000            0.000
         72              0.667            0.333             0.000            0.000
         106             0.000            1.000             0.000            0.000
         140             0.000            0.333             0.667            0.000
         174             0.000            0.667             0.333            0.000
         208             0.000            0.000             1.000            0.000
         242             0.000            0.000             0.333            0.667
         276             0.000            0.000             0.667            0.333
         310             0.000            0.000             0.000            1.000
            As an illustration, the covariates for DIM 38 can be computed as:
            F (38) = (38 − 4)/106 − 4) = 0.333 and F (38) = 1 − 0.333 = 0.667
              1                                  2
         Thus F(38) = [0.333, 0.667, 0 0]
         A random regression model can therefore be fitted as:
                       nf       nr       nr
                    i ∑
                                       +
                              +
            y   =  htd +  f b k ∑ f u jk ∑ f pe  +  e
             tijk         jtk      jtk      jtk  jk  tij
                       k=0     k=0      k=0
         where all terms are as defined in Section 9.3 but the f  is the vector of the kth spline
                                                       jtk
         function for the test day record of cow j made on day t. The same procedure described
         in Section 9.3.1 can be used in the application of the model for the analysis of data
         and interpretation of results.
         9.3.6  Random regression model for maternal traits

         Maternal genetic effects are important in growth traits in beef cattle, and models that
         account for these effects have been discussed in Chapter 7. However, the RR model
         could also be augmented to include random regressions for maternal genetic and
         maternal permanent environmental effects. Albuquerque and Meyer (2001) examined
         different orders of fit for the random regressions for both effects. One of the favoured
         models was the one in which the order of Legendre polynomials for direct genetic,
         maternal genetic, animal pe and maternal pe effects were 5, 5, 5 and 3, respectively.
            Such a model, excluding all fixed effects, could be written as:

                                      3 −
                                                 4 −
                   1−
                            2 −
                  k 1      k 1        k 1        k 1
                                    +
                 =
            y ijktd ∑  f u ji  +  ∑  f m ji ∑  f pe ji  +  ∑ f pp +  e ijktd
                                                     dtti
                               jti
                                          jti
                                                        di
                     jti
                  i=0       i=0        i=0       i=0
         where y   is the body weight of cow j taken at age t that has a dam d; u , m  and
               ijktd                                                     ji  ji
         pe  are the random regressions for direct, maternal genetic and animal pe effects for
           ji
         animal j, respectively; pp  is the random regression for dam pe effects and e   is
                               di                                            ijktd
         random error; f  and f  are the vector of the ith Legendre polynomial for body
                       jti    dti
         weight at age t for cow j and dam d, respectively. They assumed a zero covariance
         between direct and maternal genetic effects to simplify the computation. The vari-
         ance for direct effects increased from birth to 365 days while maternal genetic
         variance increased from birth to about 115 days and decreased thereafter.
          148                                                             Chapter 9
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