Page 306 - Linear Models for the Prediction of Animal Breeding Values 3rd Edition
P. 306

Taking into account contributions from animal effect (see Eqn 17.6) gives:

                 Mate is unknown              Mate is known
                                 2
                                   12
                                                                    ˆ
                 ARHS = ARHS + ( )g (uˆ )     ARHS = ARHS + (u  − 0.5u )g 12
                                                             ˆ
                      l      l   3    o            l      l   o     ma
                                                                1
                                   1
                 ARHS = ARHS − (u )( )g 12    ARHS = ARHS − (u )( )g 12
                                 ˆ
                                                             ˆ
                      l      l   l  3              l      l   l  2
            For the animal l, which is a dam with progeny having yield records, DIAG and
         ARHS from the pedigree is augmented with information from yield as:
            DIAG = DIAG + r −1
                            n
         and:
                                     ˆ
                                          ˆ
                              −1
                                              ˆ
            ARHS  = ARHS  + r (y    − hd  − b − u  − p ˆ e )
                  l       l  pa  ijklt  i  j   k    t
         for parent records and:
                                          ˆ
                                      ˆ
                              −1
            ARHS  = ARHS  + r (y    − hd  − b − 0.5(û  + û ) − p ˆ e )
                  l       l  np  ijklt  i  j      s   d     t
         for non-parent records.
            After processing all records from pedigree and yield records for the lth animal,
         the solution for the maternal effect is computed as:
             ˆ
            m  = ARHS /DIAG
              l       l     l
            The calculation of the solution for animal 5 in the first round of iteration is as follows.
         The contribution from a type 1 record in the pedigree is:
                              22
                                          12
                                                   12
            ARHS  = (mˆ  + mˆ )g  + (û  + û )g  − (û 2g )
                  5    1    2       1   2       5
                   = (0.0217 + −1.7027)0.01261 + (0 + (−1.7294))0.00336 − ((−0.5831)(2)0.0336)
                   = −0.02309
            DIAG   = (2)0.01261 = 0.02522
                 5
         The contribution from a type 2 record in the pedigree is:
            ARHS  = ARHS  + (mˆ  −  mˆ )g  + (û  −  û )g  − (û  1  g )
                                                                12
                                                      12
                                         22
                                   1
                                                 1
                  5       5     8  2  3       8  2  3       5 2
                                    1
                   = −0.02309 + (0 −   (0.4587))0.01261
                                    2
                                                             1
                     + (1.4382 −   (0.8960))0.00336 − ((−0.5831)( )0.00336)
                                1
                                2                            2
                   = −0.021675
                                22
            DIAG  = DIAG  +  g  = 0.02522 + 0.0063 = 0.03153
                             1
                 5        5  2
         The contribution from yield of progeny (animal 8) for dam 5 is:
                                     ˆ
                                          ˆ
                              −1
            ARHS  = ARHS  + r (y  − hd  − b  − u  − p ˆ e )
                                              ˆ
                  5       5   pa  8    1  1   8    5
                                 −1
                   = −0.021675 + r (40 − 30.00 − 3.679 − (1.4382) − 0)
                                 pa
                   = − 0.007724
                              −1
            DIAG  = DIAG  + r  = 0.03153 + 0.002857 = 0.034387
                 5        5   pa
         and the solution is:
             ˆ
            m  = −0.007724/0.034387 = −0.225
              5
            Solutions for permanent environmental effects are solved for after processing all
         animals for maternal effects in the current round of iteration.
          290                                                            Chapter 17
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