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Box B11  Learning and  ognition in Birds  237

  VetBooks.ir  (rather than part of their own body) dynami-  et al. 2002), suggesting that the process may
                                                      be much more flexible than would be possible
             cally to act on a goal (Bentley‐Condit and
             Smith 2010). However, it is possible that birds
                                                       It should be clear from the discussion above,
             using or even making tools habitually do so   through associative learning alone.
             using standard processes of learning, whereby   that on‐going research demonstrates that
             making  or using a  tool  of a  particular  type   many species of bird have advanced learning
             in  a  particular  way  results  in  a  reward.   and cognitive abilities, rivalling that of many
             Nevertheless, experiments suggest that habit-  mammals. As with any animal, the mixture of
             ual and non‐habitual tool users alike are able   ‘expertise’ is shaped by the ecological niche,
             to make a novel tool to solve a problem (e.g.   but if provided with the right circumstances in
             Auersperg et al. 2011, 2012; Bird and Emery   which to demonstrate their prowess, birds are
             2009; Morand‐Ferron and Quinn 2011; Weir   anything but ‘bird‐brained’.



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