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44   The Chemistry and Fertility of Soils under Tropical Weeds




                                        8000
                                     Soil Erosion (g 24 m 2 )  Erosi Tanah (g/24m2)  6000



                                        4000

                                        2000

                                          0
                                               AP    AC     CC     GA    GA-P   TS
                                                      Jenis Strip Penutup Tanah
                                                          Soil Strip Covers


                                   Fig. 3.3.  Soil erosion rates of coffee plantation in West Lampung
                                    treated with strips of weeds -  AP Arachis pintoi, AC Axonopus
                                  compressus, CC Cymbopogon citrates, GA uncut natural weeds, GA-P
                                      cut natural weeds, TS without strip (Sriyani et al., 2009).


                              As  pioneer  plants,  weeds  may  easily  adapt  to the  soil  condition  and  quickly
                           grow  and  accumulate  biomass  (Fig.  3.4).    The  production  of  high  biomass  may
                           enable  the  use  of  weeds  to  improve  the  soil  fertility.    Some  species  of  tropical
                           weeds are potential as biomass producers (Sembodo et al., 2012).  Among these
                           weeds  are  Crotalaria  lappacea,  Asystasia  gangetica,  Chromolaena  odorata,  and
                           Widelia  sp.  of  broad-leaf  weeds  and  Imperata  cylindrica,  Setaria  plicata,  and
                           Paspalum conjugatum of grasses.  However, the effect of each weed at improving
                           the soil fertility, particulary Total N, Organic C, and CEC, is different (Tabel 3.2).
                           Crotalaria lappacea and Asystasia gangetica are shown to produce high biomass
                           and increase soil organic C and CEC, while Widelia sp. significantly improves the soil
                           available P.  Therefore, the presence of weeds improved not only the soil organic
                           matter but also other soil chemical properties and also soil physical properties.
                              The presence of weeds is also reported to increase the soil enzymatic activities
                           including those of acid and alkaline phosphatases, -glucosidase, and arylsulfatases
                           (Salam et al., 2001).  Weeds also increase the soil available P (Salam et al., 2001).
                           Some weed species are also reported to be bioindicators in bioassay.  Bioassay is a
                           method  to  measure  response  of  a  particular  living  organism  to  determine  the
                           existence  or  concentration  of  a  chemical  in  a  substrate  (Satelmann,  1987).
                           Bioassay  to  detect  herbicides  usually  employs  herbicide  sensitive  plants.    In
                           addition  to  detect  the  concentration  of  herbicides  and  to  predict  the  herbicide
                           resistence, bioassay is also used to know the resistence  of weeds on herbicides.

                           Abdul Kadir Salam and Nanik Sriyani – 2019
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