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feedback loops. Negative feedback enhances stability, and over   It is difficult to understand systems fully just by focus-
                        time only those systems that are stable will persist.  ing on their individual components because systems can
                            Positive feedback loops have the opposite effect. Rather   show emergent properties, characteristics not evident in the
                        than stabilizing a system, they drive it further toward an   components  alone.  Stating  that  systems possess  emergent
                        extreme. In positive feedback, increased output leads to   properties is a lot like saying, “The whole is more than the
                        increased input, leading to further increased output. Exponen-  sum of its parts.” For example, if you were to reduce a tree
                        tial growth in a population (pp. 84–85) is one such example.   to its component parts (leaves, branches, trunk, bark, roots,
                        The more individuals there are, the more offspring can be pro-  fruit, and so on) you would not be able to predict the whole
                        duced. Another example is the spread of cancer; as cells mul-  tree’s emergent properties, which include the role the tree
                        tiply out of control, the process is self-accelerating.  plays as habitat for birds, insects, fungi, and other organisms
                            One positive feedback cycle of great concern to environmen-  (Figure 5.2). You could analyze the tree’s chloroplasts (pho-
                        tal scientists today involves the melting of glaciers and sea ice in   tosynthetic cell organelles), diagram its branch structure, and
                        the Arctic as a result of global warming (p. 516). Ice and snow,   evaluate the nutritional content of its fruit, but you would still
                        being white, reflect sunlight and keep surfaces cool. But if the cli-  be unable to understand the tree as habitat, as part of a forest
                        mate warms enough to melt the ice and snow, darker surfaces of   landscape, or as a reservoir for carbon storage.
                        land and water are exposed, and these darker surfaces absorb sun-  Systems seldom have well-defined boundaries, so decid-
                        light. This absorption warms the surface, causing further melting,   ing where one system ends and another begins can be difficult.
                        which in turn exposes more dark surface area, leading to further   Consider a smartphone. It is certainly a system—a network
                        warming (Figure 5.1b). Runaway cycles of positive feedback are
                        rare in nature, but they are common in natural systems altered by   EMERGENT PROPERTIES
                        human impact, and they can destabilize those systems.

                          FaQ        But isn’t positive feedback “good”
                                     and negative feedback “bad”?
                          Understanding negative and positive feedback in systems can
                          be difficult, because it goes against the way we use those
                          terms in everyday language. In daily life, positive feedback
                          (such as a complimentary comment on a writing assignment)   Element of  CO 2  sink     Habitat
                          is something that makes us feel good, whereas negative feed-  forest ecosystem
                          back (such as criticism on schoolwork) may make us feel bad.
                          In essence, we have been trained to view positive feedback
                          as a stabilizing force (“Keep up the good work, and you’ll
                          succeed”) and negative feedback as a destabilizing force (“You
                          need to change your approach if you’re going to succeed”).
                              In environmental systems, it’s the opposite! Nega-
                          tive feedback resists change in systems, and in doing so it                                SYSTEM
                          enhances stability, typically keeping conditions within ranges
                          beneficial to life. Positive feedback exerts destabilizing effects
                          that push conditions in systems to extremes, threatening
                          organisms adapted to the system’s normal conditions. Thus,
                          negative feedback in environmental systems typically aids liv-
                          ing things, whereas positive feedback often harms them.                  Tree


                        Systems show several defining properties                                                                  CHAPTER 5 • Envi R onm E n TA l S y STE m S   A nd E C o S y STE m E C ology
                                                                               Leaves                               Chloroplast
                        In a system stabilized by negative feedback, when pro-
                        cesses move in opposing directions at equivalent rates so
                        that their effects balance out, they are said to be in dynamic
                          equilibrium. Processes in dynamic equilibrium can con-
                        tribute to  homeostasis, the tendency of a system to main-  Acorn    Branches   Trunk        Water
                        tain constant or stable internal conditions. A system (such
                        as an organism) in homeostasis keeps its internal conditions           COMPONENTS
                        within a range that allows it to function. However, the steady   Figure 5.2 A system’s emergent properties are not evident
                        state of a homeostatic system may itself change slowly over   when we break the system down into its component parts.
                        time. For instance, Earth has experienced gradual changes in   For example, a tree serves as wildlife habitat and plays roles in
                        atmospheric composition and ocean chemistry over its long   forest ecology and global climate regulation, but you would not
                        history, yet life persists and our planet remains, by most defi-  know that from considering the tree only as a collection of leaves,
                        nitions, a homeostatic system.                       branches, and chloroplasts.                          125







           M05_WITH7428_05_SE_C05.indd   125                                                                                    12/12/14   2:56 PM
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