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Store water
                                         and regulate
                                         water flow                                Regulate
                                                                                   climate




                                        Purify                  Provide timber
                                        water                   and other       Purify
                                                       Form     resources       air           Cycle
                                                       soil
                                                                                              nutrients
                                                 Provide                                                  Provide
                                                 habitat                                                  recreation


                                            Control                                                                  Provide
                                            erosion                                                                  food
                           Pollinate
                           plants
                                                    Provide
                                                    pest             Dampen
                                                    control          impacts from
                                                                     disturbance


                                                                                                  Filter runoff
                                                                                                  and treat
                                                                                                  waste





                        Figure 5.14 Ecological processes naturally provide countless services that we call ecosystem
                          services. Our society, indeed our very survival, depends on these services.

                                                                                 Nutrients and other materials move from one  pool, or
                        Biogeochemical Cycles                                  reservoir, to another,  remaining for varying  amounts of time
                                                                             (the residence time) in each. The dinosaur, the grass, the cow,
                        Just as nitrogen and phosphorus from fertilizer on Pennsyl-  and you are each reservoirs for carbon atoms, as are sedimen-
                        vania corn fields end up in Chesapeake Bay oysters on our   tary rocks and the atmosphere. The rate at which materials move
                        dinner plates, all nutrients move through the environment in   between reservoirs is termed a flux, and the flux between any
                        intricate ways. Whereas energy enters an ecosystem from the   given reservoirs can change over time. When a reservoir releases
                        sun, flows from organism to organism, and dissipates to the   more materials than it accepts, it is called a source, and when a
                        atmosphere as heat, the physical matter of an ecosystem is   reservoir accepts more materials than it releases, it is called a
                        circulated over and over again.
                                                                             sink. Figure 5.15 illustrates these concepts in a simple manner.  CHAPTER 5 • Envi R onm E n TA l S y STE m S   A nd E C o S y STE m E C ology

                        Nutrients circulate through ecosystems                                   Reservoirs
                        in biogeochemical cycles
                                                                                                 Large flux
                        Nutrients move through ecosystems in nutrient cycles, also
                        known as biogeochemical cycles. In these pathways, chemical
                        elements or molecules travel through the atmosphere, hydro-  Source                           Sink
                        sphere, and lithosphere, and from one organism to another, in
                        dynamic equilibrium. A carbon atom in your fingernail today
                        might have helped compose the muscle of a cow a year ago,                Small flux
                        may have resided in a blade of grass a month before that, and   Short residence            Long residence
                        may have been part of a dinosaur’s tooth 100 million years   time                          time
                        ago.  After we die, the nutrients in our bodies will spread   Figure 5.15 The main components of a biogeochemical
                        widely through the environment, eventually being incorpo-  cycle are reservoirs and fluxes. A source releases more materials
                        rated by an untold number of organisms far into the future.  than it accepts, and a sink accepts more materials than it releases.  135







           M05_WITH7428_05_SE_C05.indd   135                                                                                    12/12/14   2:56 PM
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