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Intertidal zones undergo constant change

                                                                             Where the ocean meets the land, intertidal, or littoral, eco-
                                                                             systems (Figure 16.10) spread between the uppermost reach
                                                                             of the high tide and the lowest limit of the low tide. tides are
                                                                             the periodic rising and falling of the ocean’s height at a given
                                                                             location, caused by the gravitational pull of the moon and sun
                                                                             (see Figure 21.21, p. 619). High and low tides occur roughly
                                                                             6 hours apart, so intertidal organisms spend part of each day
                                                                             submerged in water, part of the day exposed to air and sun,
                                                                             and part of the day being lashed by waves. These creatures
                                                                             must also protect themselves from marine predators at high
                                                                             tide and terrestrial predators at low tide.
                                                                                 The intertidal environment is a tough place to make a liv-
                                                                             ing, but it is home to a remarkable diversity of organisms. Life
                                                                             abounds  in  the crevices  of  rocky  shorelines,  which  provide
                                                                             shelter and pools of water (tide pools) during low tides.  Sessile
                                                                             animals such as anemones, mussels, and barnacles live attached
                                                                             to rocks, filter-feeding on plankton in the water that washes
                                                                             over them. Urchins, sea slugs, chitons, and limpets eat intertidal
                                                                             algae or scrape food from the rocks. Sea stars (starfish) creep
                                                                             slowly along, preying on the filter-feeders and herbivores.
                                                                             Crabs clamber around the rocks, scavenging detritus.
                                                                                 The rocky intertidal zone is so diverse because environ-
                                                                             mental conditions such as temperature, salinity, and moisture
                         (a) Tide pools at low tide
                                                                             change dramatically from the high to the low reaches. This
                                                                             environmental variation gives rise to horizontal bands domi-
                                                                             nated by different sets of organisms arrayed according to their
                                                                             habitat needs, competitive abilities, and adaptation to   exposure.
                                                                             Sandy intertidal areas, such as those of Cape Cod, host less bio-
                                                                             diversity, yet plenty of organisms burrow into the sand at low
                                                                             tide to await the return of high tide, when they emerge to feed.




                                                                                                                 Supratidal zone
                                                                                                                 (splash zone)


                                                                                                                 Level of high tide




                                                                                                                 Intertidal zone  CHAPTER 16 • M AR in E   A nd Co A s TA l  s ys TEM s  A nd R E sou R CE s





                                                                                                                 Level of low tide

                                                                                                                 Subtidal zone



                         (b) Tidal zones
                        Figure 16.10 The rocky intertidal zone stretches along rocky shorelines between the lowest and
                        highest reaches of the tides. Fish and invertebrates inhabit tidal pools (a), and many types of algae cover
                        the rocks. The intertidal zone (b) provides niches for a diversity of organisms, including sea stars (starfish),
                        barnacles, crabs, sea anemones, corals, bryozoans, snails, limpets, chitons, mussels, nudibranchs (sea slugs),
                        and sea urchins. Areas higher on the shoreline are exposed to the air more frequently and for longer periods,
                        so organisms that tolerate exposure best specialize in the upper intertidal zone. The lower intertidal zone is
                        exposed less frequently and for shorter periods, so organisms less tolerant of exposure thrive in this zone.  445







           M16_WITH7428_05_SE_C16.indd   445                                                                                    12/12/14   3:06 PM
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