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Figure 16.11 Salt marshes occur in temperate intertidal   Figure 16.12 Mangrove forests line tropical and subtropical
                     zones where the substrate is muddy. Tidal waters flow in chan-  coastlines. Mangrove trees, with their unique roots, are adapted
                     nels called tidal creeks amid flat areas called benches, sometimes   for growing in saltwater and provide habitat for many fish, birds,
                     partially submerging the salt-adapted grasses.       crabs, and other animals.


                     Salt marshes line temperate shorelines               have developed coastal areas. Shrimp farming in particular
                                                                          has driven the conversion of large areas of mangroves. When
                     Along many of the world’s coasts at temperate latitudes, salt   mangroves are removed, coastal areas lose the ability to slow
                     marshes occur where the tides wash over gently sloping sandy   runoff, filter pollutants, and retain soil. As a result, offshore
                     or silty substrates. Rising and falling tides flow into and out of   systems such as coral reefs and eelgrass beds are more readily
                     channels called tidal creeks and at highest tide spill over onto   degraded. Moreover, mangrove forests protect coastal com-
                     elevated marsh flats (Figure 16.11). Marsh flats grow thick   munities against storm surges and tsunamis. The 2004 Indian
                     with salt-tolerant grasses, as well as rushes, shrubs, and other   Ocean tsunami (p. 60) devastated areas where mangroves had
                     herbaceous plants.                                   been removed but caused less damage where mangroves were
                        Salt marshes boast very high primary productivity and pro-  intact.
                     vide critical habitat for shorebirds, waterfowl, and many com-
                     mercially important fish and shellfish species. Salt marshes
                     also filter pollution and stabilize shorelines against storm   Fresh water meets salt water in estuaries
                     surges. However, because people desire to live and do busi-  Many salt marshes and mangrove forests occur in or near
                     ness along coasts, we have altered or destroyed vast expanses     estuaries, water bodies where rivers flow into the ocean,
                     of salt marshes to make way for coastal development. When   mixing fresh water with salt water. Estuaries are biologically
                     salt marshes are destroyed, we lose the ecosystem services   productive ecosystems that experience fluctuations in salin-
                     they provide. When Hurricane Katrina struck the Gulf Coast,   ity as tides and freshwater runoff vary daily and seasonally.
                     for instance, the flooding was made worse because vast areas   Sheltered from crashing surf, the shallow water of estuaries
                     of salt marshes had vanished during the preceding decades as   nurtures eelgrass beds and other plant life, producing abundant
                     a result of engineering of the Mississippi River, development,   food and resources. For shorebirds and for many commercially
                     and subsidence from oil and gas drilling (Chapter 15).  important shellfish species, estuaries provide critical habitat.
                                                                          For fishes such as salmon, which spawn in streams and mature
                     Mangrove forests line coasts in the tropics          in the ocean, estuaries provide a transitional zone where young
                     and subtropics                                       fish make the passage from fresh water to salt water.
                                                                             Estuaries everywhere have been affected by coastal devel-
                     In tropical and subtropical latitudes, mangrove forests replace salt   opment, water pollution, habitat alteration, and overfishing.
                     marshes along gently sloping sandy and silty coasts.  Mangroves   Think of any major estuary in the United States, from New
                     are some of the few types of trees that are salt tolerant, and they   York Harbor to Chesapeake Bay (Chapter 5) to Florida Bay to
                     have unique types of roots that curve upward like snorkels to   the mouth of the Mississippi River (Chapter 15) to San Fran-
                     attain oxygen lacking in the mud or curve downward like stilts   cisco Bay. They all have suffered pollution and other impacts
                     to support the tree in changing water levels (Figure 16.12). Fish,   from human development. Coastal ecosystems have borne the
                     shellfish, crabs, snakes, and other organisms thrive among the   brunt of human impact because two-thirds of Earth’s people
                     root networks, and birds feed and nest in the dense foliage of   choose to live within 160 km (100 mi) of the ocean.
                     these coastal forests. Besides serving as nurseries for fish and
                     shellfish that people harvest, mangroves also provide materials   Kelp forests harbor many organisms
                     that people use for food, medicine, tools, and construction.
                        From Florida to Mexico to the Philippines, half the   Along many temperate coasts, large brown algae, or  kelp,
             448     world’s mangrove forests have been destroyed as people   grow from the floor of continental shelves, reaching up toward







           M16_WITH7428_05_SE_C16.indd   448                                                                                    12/12/14   3:06 PM
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