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species—these processes and more have combined to threaten                               Biosphere
                     Earth’s biodiversity (pp. 301–308).                                                      The sum total of
                        When we look around us, it may not appear as though a                                 living things on
                     human version of an asteroid impact is taking place, but we can-                         Earth and the
                     not judge such things on the timescale of a human lifetime. On                           areas they
                                                                                                              inhabit
                     the geologic timescale, extinction over 100 years or even 10,000
                     years appears instantaneous. In contrast, speciation is a slow
                     enough process that it will take life millions of years to recover—
                     by which time our own species will most likely not be around.


                     Levels of Ecological                                                                     Ecosystem

                     Organization                                                                             A functional
                                                                                                              system consisting
                                                                                                              of a community,
                     Extinction, speciation, and other evolutionary mechanisms                                its nonliving
                     and patterns play key roles in ecology. Ecology is the scien-                            environment,
                                                                                                              and the
                     tific study of the distribution and abundance of organisms, the                          interactions
                     interactions among organisms, and the relationships between                              between them
                     organisms and their environments. It is often said that ecology
                     provides the stage on which the play of evolution unfolds. The
                     two are intertwined in many ways.

                                                                                                              Community
                     We study ecology at several levels                                                       A set of
                     Life exists in a hierarchy of levels, from atoms, molecules, and                         populations of
                                                                                                              different species
                     cells (pp. 41–47) up through the biosphere, which is the cumu-                           living together in
                     lative total of living things on Earth and the areas they inhabit.                       a particular area
                     Ecologists are scientists who study relationships at the higher
                     levels of this hierarchy (Figure 3.11), namely at the levels of the
                     organism, population, community, ecosystem, and biosphere.
                        At the level of the organism, the science of ecology describes
                     relationships between an organism and its physical environ-
                     ment. Organismal ecology helps us understand, for example,
                     what aspects of a Hawaiian honeycreeper’s  environment are                               Population
                     important to it, and why. In contrast,  population ecology exam-                         A group of
                     ines the dynamics of population change and the factors that                              individuals of a
                     affect the distribution and abundance of members of a popula-                            species that live
                                                                                                              in a particular
                     tion. It helps us understand why populations of some species                             area
                     (such as endangered honeycreepers) decline while populations
                     of others (such as ourselves) increase.
                        In ecology, a  community consists of an assemblage of
                     populations of interacting species that live in the same area.
                     A population of ‘akiapo¯la¯ ‘au, a population of koa trees, a
                     population of wood-boring grubs, and a population of ferns,
                     together with all the other interacting plant, animal, fungal,                           Organism
                     and microbial populations in the Hakalau Forest, would be                                An individual
                     considered a community. Community ecology focuses on pat-                                living thing
                     terns of species diversity and on interactions among species,
                     ranging from one-to-one interactions to complex interrela-
                     tionships involving the entire community.
                        Ecosystems encompass communities and the abiotic
                     (nonliving) material and forces with which community mem-
                     bers interact. Hakalau’s cloud-forest ecosystem consists of its
                     community plus the air, water, soil, nutrients, and energy used
                     by the community’s organisms. Ecosystem ecology reveals pat-
                     terns, such as the flow of energy and nutrients, by studying   Figure 3.11 Ecologists study questions on the levels of
                     living and nonliving components of systems in conjunction.   the organism, population, community, ecosystem, and
                     Today’s warming climate (Chapter 18) is having ecosystem-    biosphere.
               78    level consequences as it exerts impacts on the organisms of







           M03_WITH7428_05_SE_C03.indd   78                                                                                     12/12/14   2:54 PM
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