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Hakalau and many other ecosystems throughout the world.   animals in the area that it scavenges for food. For a whale,
                        As new technologies allow scientists to learn more about the   water  temperature,  salinity,  and  the  density  of  marine
                        complex dynamics of natural systems at a global scale, ecolo-  microorganisms might be critical characteristics. Each spe-
                        gists are increasingly expanding their horizons beyond eco-  cies assesses habitats differently because each species has
                        systems to the biosphere as a whole.                 different needs.
                            In the remainder of this chapter we explore ecology up   Habitat use is important in environmental science because
                        through the population level. In Chapter 4 we examine com-  the availability and quality of habitat are crucial to an organ-
                        munity ecology, and in Chapter 5 we consider ecology at the   ism’s well-being. Indeed, because habitats provide everything
                        levels of the ecosystem and biosphere.               an organism needs, including nutrition, shelter, breeding sites,
                                                                             and mates, the organism’s very survival depends on the avail-
                        Each organism has habitat needs                      ability of suitable habitats. Often this need results in conflict
                                                                             with people who want to alter or develop a habitat for their
                        At the level of the organism, each individual relates to its   own purposes.
                        environment in ways that tend to maximize its survival and
                        reproduction. One key relationship involves the specific envi-  Niche and specialization are key
                        ronment in which an organism lives, its habitat. A species’
                        habitat consists of the living and nonliving elements around it,   concepts in ecology
                        including rock, soil, leaf litter, humidity, plant life, and more.   Another way in which an organism relates to its environment is
                        The ‘akiapo¯ la¯ ‘au (Figure 3.12) lives in a habitat of cool, moist,   through its niche. A species’ niche reflects its use of resources
                        montane forest of native koa and ‘o¯ hi‘a trees, where it is high   and its functional role in a community. This includes its con-
                        enough in elevation to be safe from avian malaria.   sumption of certain foods, its role in the flow of energy and
                            Each organism thrives in certain habitats and not in oth-
                        ers, leading to nonrandom patterns of  habitat use. Mobile   matter, and its interactions with other organisms. The niche is
                                                                             a multidimensional concept, a kind of summary of everything
                        organisms actively select habitats in which to live from among   an organism does.  The pioneering ecologist Eugene Odum
                        the range of options they encounter, a process called habitat   once wrote that “habitat is the organism’s address, and the
                        selection. In the case of plants and of rooted animals (such as   niche is its profession.”
                        sea anemones in the ocean), whose young disperse and settle   Organisms vary in the breadth of their niches. Species
                        passively, patterns of habitat use result from success in some   with narrow breadth, and thus very specific requirements, are
                        habitats and failure in others.                      said to be specialists. Those with broad tolerances, able to use
                            Habitats are scale dependent. A tiny soil mite may use less
                        than a square meter of soil in its lifetime. A vulture, elephant,   a wide array of resources, are generalists. A native  Hawaiian
                                                                             honeycreeper like the ‘akiapo¯ la¯ ‘au (see Figure 3.12) is a
                        or whale, in contrast, may traverse miles upon miles of air,   specialist, because its unique bill is exquisitely adapted for
                        land, or water in just a day. Species also may have  different   feeding on grubs that tunnel through the wood of native trees.
                        habitat needs in different seasons; many migratory birds use   In contrast, the common myna (a bird introduced to Hawai‘i
                        distinct breeding, wintering, and migratory habitats.  from Asia) is a generalist; its unremarkable bill allows it to
                            The criteria by which organisms favor some habitats
                        over others can vary greatly. The soil mite may assess avail-  eat many types of foods in many types of habitats. As a result,
                                                                             the common myna has spread through virtually all areas of
                        able habitats in terms of the chemistry, moisture, and texture   the Hawaiian Islands where native birds have disappeared and
                        of the soil and the percentage and type of organic matter.   where human development has altered the landscape. Today it
                        The vulture may ignore not only soil but also topography   is one of Hawaii’s most numerous birds.
                        and vegetation, focusing solely on the abundance of dead
                                                                                 Specialists succeed over evolutionary time by being
                                                                             extremely good at the things they do, but they are vulnerable
                                                                             when conditions change and threaten the habitat or resource
                                                                             on which they have specialized. Generalists succeed by being   CHAPTER 3 • Ev ol u T i on, Bi odiv ER si T y,  A nd Po P ul AT i on E C ology
                                                                             able to live in many different places and to withstand vari-
                                                                             able conditions, but they may not thrive in any one situation
                                                                             as much as a specialist would. An organism’s habitat prefer-
                                                                             ences, niche, and degree of specialization each reflect adapta-
                                                                             tions of the species and are products of natural selection.



                                                                             Population Ecology


                                                                             Individuals of a species that inhabit a particular area make up
                        Figure 3.12 The ‘akiapo¯ la¯ ‘au lives in a habitat of cool,   a population. Species may consist of multiple populations that
                        moist, native forest on the slopes of Hawaiian volcanoes.   are geographically isolated from one another. This is the case
                        It fills a unique niche by virtue of its odd bill, whose short, straight   with Hawaii’s state bird, the ne¯ne¯ , a goose that grazes in open
                        bottom half, and long, curved top half allows it to specialize on    grassy areas (see Figure 3.1d). Originally common throughout
                        digging grubs out from native trees.                 the Hawaiian Islands, the ne¯ne¯ (pronounced “nay-nay”) was   79







           M03_WITH7428_05_SE_C03.indd   79                                                                                     12/12/14   2:54 PM
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