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             Alcids

             David A. Oehler



               Natural History

             The family Alcidae, or auk, is a group of marine, neritic, and pelagic birds with a circumpolar
             range, found exclusively in the Northern Hemisphere. The name auk was derived from the Norse
             word ãlka, given to describe the calls of these seabirds. Alcids have been divided into 11 genera
             represented by 22 species. These species are represented by six major species groups divided into
             two phyletic lines, one involving puffins and the other comprised of the auklets, murrelets, guil-
             lemots, Dovekie, and the auks. All members of this group are highly specialized, wing-propelled,
             diving birds with thickset, torpedo-shaped bodies and short wings and tails. Typical morphology
             consists of plumage that is mainly black or gray above and white below. Dramatic alterations of
             this  plumage  occur  during  the  winter  months,  with  certain  species  obtaining  gray-and-white
             plumage. The specific diving abilities of particular species, coupled with variations in prey items
             from zooplankton to fish, alleviate competition among species. There is a close correlation between
             diet to body mass and bill type (del Hoyo et al. 1996).
               Due to their prey selection, alcids are normally found in the waters of the continental shelf
             located in low and Arctic waters, with only members of the genus Synthliboramphus located in
             subtropical waters. Foraging entails remaining on the water for a majority of the year and coming
             to land only to nest and propagate. Alcids normally use islands and sea cliffs to establish breeding
             colonies of up to more than one million birds in size. Nests within these colonies usually are found
             within rocky substrates, earthen burrows, or inaccessible cavities, with only one species utilizing
             an arboreal platform nest. Adult feeding of chicks typically follows a pattern of increased provi-
             sions affected by the nutritional status of the chicks, followed by restricted amounts prior to fledg-
             ing (Hudson 1979; Ashcroft 1979; Emms 1987; Harding et al. 2002).



             ­ Criteria­for Intervention

             Very few alcid species’ chicks are encountered by rehabilitation facilities, which may be due to the
             inaccessibility of most breeding colonies and nest sites. Logging activities have resulted in a small
             number  of  Marbled  Murrelets  (Brachyramphus marmoratus)  requiring  intervention,  since  this
             species  nests  on  the  branches  of  old-growth  conifers  (Hamer  and  Nelson  1995).  Murres  and
             Razorbills (Alca torda) have an intermediate developmental strategy, allowing the chicks to depart



             Hand-Rearing Birds, Second Edition. Edited by Rebecca S. Duerr and Laurie J. Gage.
             © 2020 John Wiley & Sons, Inc. Published 2020 by John Wiley & Sons, Inc.
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