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                 Pseudochazara anthelea [Hübner, 1824] White-Banded Rockbrown


                    A species of higher altitudes, usually found above 1000
                    m. Flight is rapid, with frequent pauses to alight on
                    rocks in open, mountainous areas. P. anthelea shows
                    marked sexual dimorphism; the upperside ground
 #                                                                                                                                                                                      © Christodoulos Makris    © Dubi Benyamini    © Dubi Benyamini
                                                                                                                                                                                                              © Christodoulos Makris
                    colour of the smaller male is dark brown, verging on
                    black, relieved by a wide white postdiscal band on                                                                                    DGXOW
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                    the fore- and hindwings. Male forewing underside                                                                                      HJJ  1  1    2    3    4    5    6    7    8    9   10  11  12
                                                                                                                                                     lar v a
                                                                                                                                                 pupa
                    ground colour is white with two prominent ocelli,
                    visible also on the upperside.  The female’s upperside
 21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 7 - A | 22-01-02 | 12:34:17 | SR:-- | Magenta
 21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 7 - A | 22-01-02 | 12:34:17 | SR:-- | Yellow
                    ground colour is brown, with a wide post-discal band
 #21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 7 - A | 22-01-02 | 12:34:17 | SR:-- | Black
 21603-BRACHA-PARPAR - 21603-BRACHA-PARPAR | 7 - A | 22-01-02 | 12:34:17 | SR:-- | Cyan
                    of orange on the forewing; within this band are two                                                                                                                            © Dubi Benyamini            © Dubi Benyamini
                    large ocelli, visible also on the underside. Forewing
                    underside markings are similar to the male, but with                          DGXOW  1  1    2    3    4    5    6    7    8    9   10  11  12
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                    ground colour orange. Hindwing underside is mottled                 lar v a
                                                                             pupa
                    grey/brown. Males eclosed usually few days before the
                    female (Hesselbarth et al., 1995(2): 961). Males are
                    territorial and often found on steep rocky slopes - in
                    Cyprus usually above 1000 m (Wakeham-Dawson et                                                                                                                                                          © Martin Gascoigne-Pees
                    al., 2003). Overlaps with the similarly marked (female                                                                                                            Subterranean pupal chamber
                    only) P. telephassa in the Hatay region of S Turkey –
                    see below. The females (and possibly males) aestivate                                                                                                                          Ļ
                    during the summer months and resume activity for egg
                    laying in autumn. In courting the male land next to the
                    female, position itself in front of her, open and tilt its
                    wings toward the female, go around the female trying to                                                                                                                         © Dubi Benyamini           © Dubi Benyamini
                    excess her genitalia, the female remained motionless
                    with her abdomen hide between her close wings. After
                    several minutes the male gave up and flew away (DB
                    pers. obs.). In Cyprus adult's preferred nectar sources                                                                                                                      Ļ  last larval
                    are the purple flowers of  Centaurea aegilophila and                                                                                                                           exuviae
                    Ptilostemon  chamaepeuce (Asteraceae) (Makris
                    2003: 250 & pers. comm. to DB). In S Turkey anthelea
                    ZDV UHFRUGHG RQO\ RQFH LQ ùVNHQGHUXQ  +DWD\  $WDKDQ
                    et al., 2018: 83, and pers. comm.). Possibly extinct in
                    Lebanon since its sole record by Lederer near Beirut
                    (1857).                                                                                                                                                              Ļ                   © Dubi Benyamini
                                                                                                                                                                         © Dubi Benyamini  Subterranean pupal chamber          © Dubi Benyamini
                    %LRORJ\
                    )OLJKW SHULRG  April to September, 250 to 1900 m in
                    Cyprus (Makris 2003: 250). The single record from S.
                    Turkey is from May at 1400 m (Atahan  et al., 2018:
                    83).
                                                                                                              © Eddie John
                    /LIH  KLVWRU\  univoltine. Laid egg is white, barrel
                    shape, 1 mm high and 0.5-0.6 mm max diam. with 20-  'LVWULEXWLRQ
                    27 vertical ribs. Emerging L1 consumes the egg shell,
                    is 1.5-2 mm long, light brown with darker brown dots   TL: Asia Minor. Ranges across Albania, Macedonia, Bulgaria,
                    along the body and head. Single known L5 was found   Greece, Turkey (rare in Hatay), Cyprus (absent from other                                                                   © David Jutzeler           © David Jutzeler
                    under a stone and photographed by Wolfgang Wagner   countries of the Levant) and N Iraq. Rebel (1916) described
                    in Ida Mt., C Crete, ~30 Km SW Heraklion; chunky ca   the Cyprus P. anthelea as ‘Satyrus anthelea (Hb.) acamanthis
                    26-28 mm long larva is greyish-brown with longitudinal   N subsp.’ Van Oorschot et al., (1987) reluctantly designated
                    whitish broad strips along all its body, mid-dorsal and   a lectotype for  P.  anthelea  acamanthis, accepting some
                    side darker broken lines and sub-dorsal line of arches,   of the distinguishing morphological characters noted
                    the short tail is off-white (illust.). Overwintering in the   by Rebel, but the ssp. status was questioned by Olivier
                    larval stage. The pupa is 15-16 mm long and 6-6.5 mm   (1996), who synonymised P. anthelea acamanthis with P.
                    wide, is radish-brown, smooth and resembles that of   anthelea anthelea. Subsequent work by Wakeham-Dawson
                    P. pelopea – illust. The adult eclosed after 3-4 weeks.   et al., (2003) supported Olivier’s view, but proposed
                    (Wolfgang Wagner & Marios Aristophanous pers.    further investigation. Here, with the subspecific status of P.
                    comms. to DB).                                   anthelea acamanthis at best only weakly defined, we follow
                                                                     Olivier and Wakeham-Dawson et al., (op. cit.) and lump the
                    5HFRUGHG KRVWSODQWV  Poaceae (Gramineae) spp.    Cyprus taxon with the nominotypical subspecies.                                                     © Dubi Benyamini            © David Jutzeler           © David Jutzeler
              108                                                                                                                                                                                                                        101
           2Butterflies of the Levant danaidae satyridae.indd   108                                            12/30/2021   4:25:13 PM  2Butterflies of the Levant danaidae satyridae.indd   101                                   12/30/2021   4:24:54 PM
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