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Both species prefer dry habitats in Africa and
Arabia and their hostplants in the families
Euphorbiaceae and Solanaceae? are common
on both sides of the Red Sea mountain chains
that are practically unexplored.
Very limited recent exploration of N Red Sea
mountain ranges (Benyamini & Müller, 2020)
resulted in discovery of over ten regional new
species, expending northwards due to climate
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1 2 3 4 5 6 7 8 9 10 11 12
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1
1
10
1
DGXOW
HJJ 12 3 3 4 5 91 01 1 1 2
change. Tropical storms moving northwards e.g. lar v a © Dubi Benyamini
pupa
RST that serves as conveyers for African and 10
Arabian butterflies are carrying and pushing 1 1 1 1 1 1
the butterflies northwards giving hope for
additional relocated species including one or
both these Jokers. Migrating Byblia (anvatara)
acheloia (Larsen, 1978; Larsen & Larsen,
1980: 50) will possibly be the first to penetrate
S Levant.

© Nancy van der Poorten
%\EOLD LOLWK\LD 8
A local and rare species to the southern
outskirts of the Levant. In Dhofar S Arabia
it prefers dense vegetation while in W Africa
it is found only in very dry areas. Adults are
attracted to flowers of Heliotropium spp. and
sap of damaged Solanum incanum (Larsen &
Larsen, 1980: 50; Larsen, 1982: 33).
S Sinai massif, Wadi El-Arbain, 1600 m, 5.11.1974
overwintering specimen

© Leah Benyamini
2 2 2 2 2

© Dubi Benyamini
© Bart Wursten
%\EOLD DQYDWDUD
A rare species in the southern margins of the
Levant. Seasonally dimorphic species, with
specimens of the dry season having much © Dubi Benyamini © Dubi Benyamini
darker underside. Quite “common in African
shrublands where it flies close to the ground
© Dubi Benyamini 32), but in Asir, SW Arabia (as B. acheloia © Dubi Benyamini © Dubi Benyamini © Ofir Tomer © Benny Shalmon
and often inside bushes” (Larsen, 1982:

Wallengren, 1857) is found in low vegetation
“underneath large trees and shrubs on slopes
and riverbanks” (Pittaway, 1985).

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2Butterflies of the Levant danaidae satyridae.indd 148 12/30/2021 4:27:20 PM 2Butterflies of the Levant danaidae satyridae.indd 157 12/30/2021 4:27:42 PM