Page 48 - Linear Models for the Prediction of Animal Breeding Values 3rd Edition
P. 48

-1
         Without inbreeding, d =  16  .
                            ii  11
            If only the maternal grandsire (k) of animal i is known, add:
             −1
            d   to the (i,i) element
             ii
              −1
            −d /4 to the (k,i) and (i,k) elements
               ii
             −1
            d /16 to the (k,k) element
             ii
         Without inbreeding, d -1  =  16  .
                            ii  15
            If only the sire (s) of animal i is known, add:
             −1
            d  to the (i,i) element
             ii
              −1
            −d /2 to the (s,i) and (i,s) elements
               ii
             −1
            d /4 to the (s,s) element
             ii
                                4
         Without inbreeding, d -1  =  in this situation, as in Section 2.4.1.
                            ii  3
            When s and k are unknown, add:
             −1
            d   to the (i,i) element
             ii
              −1
         and d  = 1.
              ii
         2.6 An Example of the Inverse of a Sire and Maternal Grandsire
         Relationsip Matrix
         A pedigree consisting of sires and maternal grandsires set up from the pedigree in
         Table 2.1 is:
                         Sire   Sire of sire  Maternal grandsire of sire
                         1      Unknown      Unknown
                         4      1            Unknown
                         5      4            1

         Recoding sires 1 to n, the pedigree becomes:

                                  1     Unknown  Unknown
                                  2     1        Unknown
                                  3     2        1
                                  ⎡ 1.0 0.5   0.5  ⎤
                                  ⎢                ⎥
         Using Eqns 2.6 and 2.7, A =  0.5 1.0  0.625
                                  ⎢                ⎥
                                  ⎢ ⎣ 0.5 0.625 1.125⎥ ⎦
         Note that the relationship among sires is the same as in A calculated from the full
         pedigree in Section 2.2.
                                             ⎡  1.0   0.0 0.0⎤
              −1
         The T  matrix for the pedigree is: T −1  =  ⎢ ⎢ −0.5  1.0 0.0 ⎥ ⎥
                                             ⎢ ⎣ −0.25  −0.5 1.0⎥ ⎦


          32                                                              Chapter 2
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