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(a) Tropical dry forest                    400        (a) Savanna                               300

                           70
                          Mean monthly temperature (ºC)  50        100  Mean monthly precipitation (mm)  Mean monthly temperature (ºC)  40  80 Mean monthly precipitation (mm)
                                                                                                                        200
                                                                   300
                           60
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                                                                   200
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                           40
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                            0
                               Jul Aug Sep Oct Nov Dec Jan Feb Mar Apr May Jun
                                    Time of year (Southern Hemisphere)  0         0 Jul Aug Sep Oct Nov Dec Jan Feb Mar Apr May Jun 0
                                                                                        Time of year (Southern Hemisphere)
                        (b) Darwin, Australia                                 (b) Harare, Zimbabwe
                         Figure 4.23 Tropical dry forests experience significant   Figure 4.24 Savannas are grasslands with clusters of trees.
                         seasonal variation in precipitation and relatively stable,   They experience slight seasonal variation in temperature but
                         warm temperatures. Climatograph adapted from Breckle, S.W., 2002.  significant variation in rainfall. Climatograph adapted from Breckle,
                                                                             S.W., 2002.
                            Despite  this  profusion  of  life,  tropical  rainforests  have
                        poor, acidic soils that are low in organic matter. Nearly all nutri-  Savanna    Drier tropical regions give rise to  savanna
                        ents present in this biome are contained in the plants, not in the   (Figure 4.24), tropical grassland interspersed with clusters
                        soil. An unfortunate consequence is that once tropical rainfor-  of acacias or other trees. The savanna biome is found across
                        ests are cleared, the nutrient-poor soil can support agriculture   stretches  of Africa,  South America, Australia,  India,  and
                        for only a short time (p. 239). As a result, farmed areas are   other dry tropical regions. Precipitation in savannas usually
                        abandoned quickly, and farmers move on and clear more forest.  arrives during distinct rainy seasons, whereas in the dry sea-
                                                                             son grazing animals concentrate near widely spaced water
                        Tropical dry forest    Tropical areas that are warm year-  holes. Common herbivores on the African savanna include   CHAPTER 4 • S PEC i ES   i n TERA CT i on S   A nd Co mmuni T y E C ology
                        round but where rainfall is lower overall and highly seasonal   zebras, gazelles, and giraffes. Predators of these grazers
                        give rise to tropical dry forest, or tropical deciduous forest   include lions, hyenas, and other highly mobile carnivores.
                        (Figure 4.23), a biome widespread in India,  Africa, South
                        America, and northern Australia. Wet and dry seasons each   Desert    Where rainfall is very sparse, desert (Figure 4.25)
                        span about half a year in tropical dry forest. Organisms that   forms. The driest biome on Earth, most deserts receive well
                        inhabit tropical dry forest have adapted to seasonal fluctua-  under  25 cm (9.8  in.) of  precipitation  per year, much of  it
                        tions  in  precipitation  and  temperature.  For  instance,  many   during isolated storms months or years apart. Some deserts,
                        plants are deciduous and leaf out and grow profusely with the   such as Africa’s Sahara and Namib deserts, are mostly bare
                        rains, then drop their leaves during dry times of year.  sand dunes; others, such as the Sonoran Desert of Arizona and
                            Rains during the wet season can be heavy and, coupled   northwest Mexico, receive more rain and are more heavily
                        with erosion-prone soils, can lead to severe soil loss where   vegetated.
                        people have cleared forest. Across the globe, we have con-  Deserts are not always hot; the high desert of the west-
                        verted a great deal of tropical dry forest to agriculture. Clearing   ern United States is positively cold in winter. Because deserts
                        for farming or ranching is straightforward because vegetation   have low humidity and little vegetation to insulate them from
                        is lower and canopies less dense than in tropical rainforest.  temperature  extremes, sunlight readily  heats them in the   115







           M04_WITH7428_05_SE_C04.indd   115                                                                                    12/12/14   2:55 PM
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