Page 387 - Anatomy and Physiology of Farm Animals, 8th Edition
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372 / Anatomy and Physiology of Farm Animals

          tongue, where the digestive tract continues   (A)
          as the pharynx.
  VetBooks.ir    comprises the hard palate rostrally and the
            The  dorsal  wall  of  the  oral  cavity

          soft  palate caudally. The hard palate is
          formed  by  horizontal  elements of  the
            incisive, maxillary, and palatine bones, and
          its thick mucous membrane covering is
          characterized by prominent transverse folds
          called palatine rugae. The soft  palate is a
          musculomucosal sheet that extends toward
          the base of the epiglottis (see Chapter 19).
          The equine soft palate is exceptionally long,
          and the horse is uniquely unable to actively
          lift the soft palate so as to permit passage of   (B)
          air from the oral cavity to the larynx. For this
          reason, horses are obligate nose breathers,
          unable to breathe through the mouth unless
          the soft palate is abnormally displaced dor­
          sad from its  normal position ventral to the
          epiglottis (Fig. 20‐2).


          Teeth

          Teeth are arranged in two dental arcades,
          one associated with the mandible and one
          with  the  incisive  and  maxillary bones.
          Farm animals typically have a gap in each
          arch between the front teeth (incisors) and   Figure  20-2.  (A) The equine soft palate nor­
          the cheek teeth (see Fig. 4‐4); such a physi­  mally lies ventral to the margins of the epiglottis,
          ologic gap is a diastema. The bit of the bri­  except during swallowing when it is briefly lifted
                                                  by the passage of the food/water being swallowed.
          dle lies in the horse’s diastema.       This position provides a broad, unobstructed pas­
            Mammals typically exhibit heterodonty.   sageway for air. (B) With “dorsal displacement of
          That is, they have various types of teeth that   the soft palate,” the soft palate is inappropriately
          are specialized for different aspects of   lifted to the dorsal side of the epiglottis. The long
            prehension and mastication. All domestic   palate hangs into the laryngeal vestibule and con­
          animals also are  diphyodont. This means   stitutes a partial obstruction to air movement. In
          they develop a set of deciduous teeth (also   this position, the horse may breathe through his
          called baby teeth or milk teeth) that fall out   mouth, a pattern not normally possible.
          and are replaced with permanent teeth. As
          growing teeth emerge from the gums, they   examination of the wear patterns of the
          are said to  erupt. When their occlusal   occlusal surfaces.
          (grinding) surfaces meet those of the teeth   A tooth is anchored by its  root in a
          in the opposing arcade, they are said to have   socket of bone called an  alveolus.
          come into wear. Eruption times of teeth are   Connective tissue, the  periodontium
          consistent enough to permit accurate aging   (also called periodontal membrane),
          of young animals by observing their denti­  firmly attaches the root to the surround­
          tion (see Tables 20‐1 and 20‐2). The age of   ing bone in a specialized joint, the gom-
          animals with a full set of permanent teeth   phosis (see Chapter 6). The crown is the
          can be only roughly estimated through   part of the tooth visible above the mucous
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