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          (A)                                   (B)
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          Figure 20-8.  (A) Fungiform papillae from rabbit tongue (4×). Notice the presence of taste buds on the
          lateral sides of papillae (circle). (B) Close up of taste buds (40×). Arrowhead indicates pore at apex of taste
          bud through which dissolved substances can reach the taste cells of the bud. Source: photomicrographs
          courtesy of Gretchen Delcambre, Colorado State University, Colorado, USA.

          which is thickly covered with prominent    Care must be exercised not to enter this
          conical papillae. Similar cornified projec­  diverticulum while passing a stomach tube
          tions cover the inside of the lips and cheeks.  or giving medications via a balling gun.

                                                  Tonsils
          Pharynx
                                                  Tonsils  are more or  less circumscribed
          The pharynx (pl. pharynges) is the com­  aggregations of lymphatic nodules found
          mon passage for food and air, is caudal to   in association with the mucous membranes
          oral and nasal cavities, and is lined by   of the mouth and pharyngeal region
          mucous membrane and surrounded by       (Fig. 20‐10). The histologic organization of
          muscles. The pharynx can be divided into   tonsils is described in Chapter 16.
          nasal (nasopharynx), oral (oropharynx),    In humans, each of two obvious pala-
          and laryngeal (laryngopharynx) portions,   tine tonsils lies in a pocket on the lateral
          so named for their association with these   wall of the pharynx ventral to the soft pal­
          regions (see Fig. 19‐2).                ate and adjacent to the base of the tongue
            The muscles of the walls of the pharynx   (these are readily  seen  with a  mirror as
          are responsible for the orderly directing of   elongate  nodules  on each side of your
          air, food, and liquids in such a way that air   throat, unless, of course, they have been
          from the nasal cavity is directed into the lar­  removed). In the horse, ox, and sheep,
          ynx and food and liquids are directed into the   these palatine tonsils are in about the same
          esophagus. Thus, the paths of air and swal­  relative position, within the submucosa,
          lowed substances must cross in the pharynx;   and completely covered by mucous mem­
          pharyngeal dysfunction can have severe   brane. In other words, the tonsils do not
            consequences for the airway, which must be   project into the pharynx at all in these ani­
          protected from foodstuffs (Fig. 20‐9).  mals. The ruminant palatine tonsil features
            The pharyngeal recess in the horse is a   a deep cleft, the  tonsillar crypt, which
          median niche at the caudodorsal angle of   increases  the  surface area through which
          the nasopharynx (Fig. 20‐10). The pig has a   the lymphatic tissue may come in contact
          pharyngeal diverticulum that opens into   with antigens (Figs. 19‐2 and 20‐7). In the
          the dorsal wall of the pharynx near the   pig the palatine tonsils lie within the sub­
          beginning of the esophagus.             stance of the soft palate.
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