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            passes and the function of any of those   kidney  and  the distinctively  lobated  kid-
            special vascular structures.
                                                  neys of the ox (Fig. 23‐1A).
  VetBooks.ir    • What is a countercurrent exchange   abdominal cavity on each side of the aorta
                                                     The kidneys are in the dorsal part of the
            mechanism and how does this contrib-
            ute to urine formation?               and caudal vena cava, just ventral to the
             • Briefly describe how aldosterone, the   first few lumbar vertebrae. In most domes-
            renin–angiotensin system, and antidiu-  tic animals, the right kidney is slightly
            retic hormone regulate osmolality and   more cranial than the left, with the cranial
            fluid balance.                        pole of the right kidney lying snugly in a
             • What is the role of carbonic anhydrase   complementary fossa of the liver. The left
            in the body and where is it found?    kidney tends to be more pendulous, and in
             • How is sodium and potassium concen-  ruminants, the forestomach may push the
            tration regulated in the body?        left kidney to the right as far as the median
             • Describe the major buffering systems   plane or beyond, particularly when the
            for maintaining acid‐base balance within   rumen is full. The kidneys are described as
            the body.                             being retroperitoneal in location, reflect-
             • Compare and contrast the role of the   ing their position outside the peritoneal
            kidney versus lung in maintaining acid‐  cavity where they are more closely attached
            base balance.                         to the abdominal wall by fascia, vessels,
             • Define the major components of each of   and  peritoneum  than  are  most  other
            the primary metabolic or respiratory   abdominal organs. A tough connective tis-
            acid‐base imbalances.                 sue capsule surrounds the entire kidney.
                                                     The medial aspect of each kidney fea-
                                                  tures a concavity, the hilus, where arteries
             he urinary system consists of two    and nerves enter the kidney, and the ureter,
          T kidneys, two ureters, the urinary blad-  veins, and lymphatic vessels leave. The
          der, and the urethra. The paired kidneys   wide origin of the ureter in the kidney is
          remove waste products from the blood,   the renal pelvis. The renal pelvis receives
          help regulate the composition of plasma,   urine from the  collecting tubules of the
          and perform certain hormonal functions.   kidney. The  bovine kidney lacks a renal
          The system of tubules in each kidney coa-  pelvis, the ureter instead arising directly
          lesces  into  a  single  mucomuscular  tube,   from the coalescence of individual calyces
          the ureter, which extends caudad to empty   (discussed later).
          into the urinary bladder, a distensible res-  The portion of the kidney immediately
          ervoir for the storage of urine. When full,   surrounding the renal pelvis is the renal
          the  urinary bladder  discharges  the  urine   medulla, which appears striated because of
          through the urethra to the outside of   the radially arranged collecting tubules. In
          the body.                               addition to collecting tubules, the medulla
                                                  also contains some loops of Henle (descend-
                                                  ing and ascending loops). The medulla is
          Anatomy of the Kidney                   surmounted peripherally by the renal cortex,
                                                  in which reside the  renal corpuscles, the
          The kidneys are paired reddish‐brown    histological units  of  filtration. The  cortex
          organs that filter plasma and plasma con-  has a granular appearance because of the
          stituents from the blood and then selec-  large number of these renal corpuscles; also
          tively reabsorb water and useful constituents   found in the cortex are proximal and distal
          from the filtrate, ultimately excreting   convoluted tubules and other segments of
          excesses and plasma waste products. The   loops of Henle (discussed later).
          kidneys of most animals are roughly bean‐  The medulla and cortex are arranged in
          shaped, with the exceptions among domes-  units called lobes, cone‐shaped aggregates
          tic animals of the heart‐shaped right equine   of renal tissue. The medullary portion of
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