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Digestive system (apparatus digestorius)   193




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                  10.18  Floor of the mouth with mucous glands (chicken).   10.19  Pharyngeal glands (chicken). Azan stain (x40).
                  Azan stain (x40).



                    lary gland (glandula maxillaris) is located rostrally in   Tubular digestive organs
                    the roof of the mouth. It empties via a duct caudal to   The tubular digestive organs can be divided into cranial
                    the bill tip organ of the upper beak. An abundance of   (oesophagus and stomach), middle (duodenum, jejunum,
                    taste buds surrounds the duct orifice. At the angle of the   ileum) and caudal (caecum, colon, rectum) portions and
                    mouth, the glandula angularis oris is similarly drained   the anal canal (Figure 10.1).
                    by a single duct. The palate contains the openings of
                    multiple ducts of the lateral and medial palatine glands   Structure of tubular digestive organs
                    (glandulae palatinae). Numerous taste buds surround   Except for regional, functionally determined specialisa-
                    these orifices. The secretions of the mandibular glands   tions, the wall of tubular digestive organs is consistent in
                    (glandulae mandibulares) (Figure 10.18) and lingual   its basic structure (Figure 10.20). From interior to exterior,
                    glands (glandulae linguales) pass through many ducts   it is composed of the following layers:
                    to the floor of the mouth. Additional glands, located   ·  tunica mucosa:
                    more caudally, empty into the oropharynx via several     − epithelium (epithelium mucosae),
                    ducts (Figure 10.19).
                                                                        − lamina propria (lamina propria mucosae),
                                                                        − muscular lamina (lamina muscularis mucosae),
                  Pharynx                                          •  tela submucosa,
                  Due to its relationship to the nasal and oral cavities,   ·  tunica muscularis:
                  the epithelium of the pharynx varies with region. In     − circular layer (stratum circulare),
                  the nasopharynx, the  pharyngeal  mucosa  is lined by     − longitudinal layer (stratum longitudinale) and
                  pseudostratified ciliated epithelium containing gob-  •  tunica adventitia.
                  let cells. Serous and mixed glands are present in the
                  propria-submucosa.
                     The epithelium of the oropharyngeal mucosa is strati-  Within body cavities, the following layers are present in
                  fied squamous. The lamina propria encloses lymphoid tissue   place of a tunica adventitia:
                  that varies with species in its degree of organisation (lym-  ·  tela subserosa,
                  phoid follicles, tonsils) (see Veterinary Anatomy of  Domestic   ·  tunica serosa:
                  Mammals: Textbook and Colour Atlas). The connective tissue     − lamina propria serosae and
                  layer is underlaid by elastic fibres and merges indistinctly     − epithelium (mesothelium serosae).
                  with a tela submucosa containing mucous glands.
                     External to the pharyngeal submucosa is an inner fas-  These layers form a musculomembranous tube that also
                  cial layer, a layer of striated muscle, an outer fascial layer  occurs in similar form elsewhere in the body (e.g. urinary
                  and a tunica adventitia.                       tract).









       Vet Histology.indb   193                                                                                  16/07/2019   15:00
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