Page 260 - Anatomy and Physiology of Farm Animals, 8th Edition
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Sense Organs / 245

                  Animals  with  injury  to  one  side  of   open into a shallow furrow near the
                                                        mucocutaneous junction of the eyelid. The
  VetBooks.ir  their vestibular system express the      openings of their ducts are readily seen
               vestibular reflexes in the absence of
                                                        when the margin of the lid is everted, and
               appropriate stimuli. These individuals
               commonly have resting nystagmus, a       the glands themselves are usually visible
               head tilt, and a tendency to lean, circle,   as  yellowish‐white columns under the
               or fall to one side.                     mucous membrane on the inner surface
                                                        of  the  lid.  The  tarsal  glands  produce  an
                                                        important oil layer of the tear film.
               Vision                                      The conjunctiva is the mucous mem­
                                                        brane that lines the inside of the palpebrae
               The eye is an elaborate organ whose      and part of the anterior surface of the
                 primary function is to collect and focus   eyeball (excluding the clear cornea). Diseases
               light upon the photosensitive retina. It   of the conjunctiva are the most common
               lies within a coneshaped cavity of the skull,   eye  disorders.  The  conjunctiva  is  rich  in
               the orbit, which houses the eyeball (globe)   mucus‐producing glands, lymphocytes,
               and a number of other soft tissue struc­  nerves, and blood vessels. The very small
               tures, the  ocular  adnexa (e.g., muscles,   space between the eyelids and the surface
               glands), that act upon the eyeball in service   of the eye is the conjunctival sac, and it is
               of its light‐collecting function. Unlike the   into this space that eye medications are
               human orbit, which is a complete bony    usually instilled.
               cone, the ventral portion of the orbit of   Inflammation of the conjunctiva, or
               domestic species is bounded by soft tissues,   conjunctivitis, is characterized by red-
               notably the pterygoid muscles.           ness and increased mucus production.
                                                        Cancer eye, squamous cell carcinoma of
                                                        the conjunctiva, is relatively common in
               Ocular Adnexa                            Hereford, Holstein, and shorthorn
                                                        breeds of cattle.
               Palpebrae.  Two  mobile  folds  of  haired   Domestic species possess a nictitating
               skin protect the anterior aspect of the   membrane, or third eyelid. This is a fold
               eyeball. These are the eyelids, or       of mucous membrane arising from the
               palpebrae. The gap between the margins   ventromedial aspect of the conjunctival
               of the two palpebrae is the  palpebral   sac between the eyeball and palpebrae. It is
               fissure, and its size is controlled by the   given rigidity by a T‐shaped cartilage
               muscles  of  the  eyelids.  A  layer  of  dense   within it, and it smooths the tear film and
               connective tissue in the palpebrae, the   protects the cornea.
               tarsal  plate (tarsus), affords them a      The third eyelid has at its base a serous
               certain amount of rigidity. The palpebrae   gland, called simply the gland of the third
               have  thin  but  otherwise  typical  haired   eyelid, which normally contributes about
               skin on their superficial side. This skin   50% of the tear film. Ruminants, the pig,
               ends  at  the  lid  margin  with  a  sharp   and laboratory rodents possess another
               transition to mucous membrane. The       deeper gland associated with the third
               margin may have cilia (eyelashes).       eyelid. This is the harderian gland. It, too,
                  Abundant modified sweat and sebaceous   contributes to the tear film.
               glands are associated with the lid margin.
               A row of large modified sebaceous glands,   Lacrimal Apparatus.  The  lacrimal
               the  tarsal glands (meibomian glands),   apparatus comprises a series of serous,
               is especially clinically significant, as they   seromucoid, and  mucous  glands  and  the
               commonly become infected, develop        duct systems that drain their secretions
               tumors, or become impacted. The tarsal   from the conjunctival sac (Fig.  12‐11).
               glands are present in both palpebrae and   It  provides a moist environment for the
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