Page 266 - Anatomy and Physiology of Farm Animals, 8th Edition
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               Visual Field and Light Path              where objects are simultaneously viewed by
                                                        both eyes, is the binocular field; the visual
  VetBooks.ir  The part of the environment from which   cortex evaluates the slightly different view

                                                        from each eye and uses the information to
               light will enter the eyes and stimulate the
               retinas is the visual field (Fig. 12‐15). In   provide depth perception. Prey animals, on
               predators and arboreal animals, such as   the other hand, have lateral eyes with a much
               birds and primates (for whom accurate    smaller binocular field. Such eye placement
               depth perception is essential), the eyes are   increases the peripheral vision so that the
               placed so that the visual fields overlap to   combined visual field is  nearly completely
               varying degrees. This region of overlap,   panoramic. Such vision is monocular (seen
                                                        only with one eye) and therefore lacks very
                                                        accurate depth cues, but the clear advantage
                                                        of this wide field of view for a prey animal
                                                        needs no explanation.
                                                           Light traveling from the visual field to
                            Binocular field
                              (both eyes)               the retina passes through a series of trans­
                                                        parent media that refract and focus it on
                                                        the light‐sensitive retina of the posterior
                                                        part of the globe. These  dioptric media
                                                        include the cornea, the aqueous, the lens,
                                                        and the vitreous body. As indicated before,
                                                        the cornea is actually the most refractive
                                                        medium of the eye, but the lens is the only
                                                        part of the light path with the ability to
                                                        change its refractive index. This property
                                                        makes it the organ of accommodation for
                                                        focus on near objects. Light entering the
                Monocular field        Monocular field  vitreous chamber of the eye is bent by the
                   (left eye)            (right eye)
                                                        more  anterior  parts  of  the  eye  in  such  a
                                                        way that the image that is focused on the
                              Blind area
                                                        retina is inverted and reversed.
                                                           The site where the ganglion cell axons
                                                        leave the eye (the optic disk) has no pho­
                                                        toreceptors and is therefore considered
                                                        the blind spot of the retina. In primates, a
                                                        region dorsolateral to the optic disk is
                                                        relatively free from large blood vessels
                                                        and especially densely packed with photo­
                                                        receptors (particularly cones). This is the
                                                        region of greatest visual acuity, the macula.
                                                        Domestic animals lack a macula.


                                                        Visual Pathways of the Brain
               Figure 12-15.  Visual fields of the ox as viewed
               from above. The region seen by both eyes (binoc­
               ular field) is the region of best depth perception,   The electrical information generated by
               but the nose creates a wedge‐shaped blind spot   exposure of photoreceptors to light under­
               directly in front of the animal. Peripheral vision   goes initial neural processing within the
               (the monocular fields) creates a nearly complete   retina. This information ultimately leaves
               circle  of  vision  around  the  animal,  except  for  a   the eye via the optic nerve, the fibers of
               small region directly behind.            which are the axons of the ganglion cells of
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